Entry
Name
pyruvate carboxylase;
pyruvic carboxylase
Class
Ligases;
Forming carbon-carbon bonds;
Ligases that form carbon-carbon bonds (only sub-subclass identified to date)
BRITE hierarchy
Sysname
pyruvate:carbon-dioxide ligase (ADP-forming)
Reaction(IUBMB)
ATP + pyruvate + HCO3- = ADP + phosphate + oxaloacetate [RN:
R00344 ]
Reaction(KEGG)
Substrate
Product
Comment
A biotinyl-protein containing manganese (animal tissues) or zinc (yeast). The animal enzyme requires acetyl-CoA.
History
EC 6.4.1.1 created 1961
Pathway
ec00720 Carbon fixation pathways in prokaryotes
ec01120 Microbial metabolism in diverse environments
Orthology
K01959 pyruvate carboxylase subunit A
K01960 pyruvate carboxylase subunit B
Genes
PLEU : 114685980 114709141
XLA : 108715122 398811(pc.1.L)
DRE : 553283(pcxa) 58068(pcxb)
SRX : 107712165 107715262 107721657 107756168
SANH : 107661088 107661964 107669309 107699965
SGH : 107548233 107559404 107575801 107583287
CAUA : 113038514 113064144 113106607
PPRM : 120477636(pcxb) 120494698(pcxa)
MAMB : 125253542(pcxa) 125265630(pcxb)
IPU : 108267638(pcxb) 108278882(pcxa)
PHYP : 113540156 113541475
SMEO : 124386938(pcxb) 124398036(pcxa)
TFD : 113640317(pcxb) 113662088(pcxa)
AMEX : 103022961 103039377
EEE : 113573494(pc) 113586026
SASA : 106563498 106603157
STRU : 115154661 115204260
OTW : 112228804(pcxa) 112231340
OMY : 110509959 110518778 110533971
OGO : 124007431 124007758 124008783
SALP : 111950242 111964167
CCLU : 121541827 121554136 121579479
PKI : 111835210(pc) 111839535
AANG : 118224356 118235245
ARUT : 117398547 117967856
DSI : Dsimw501_GD10044(Dsim_GD10044)
DMO : Dmoj_GI20741 Dmoj_GI20742
AALB : 109397766 109428793 115269413
CCAL : 108623652 108633100
PPYR : 116168283 116168912
TNL : 113503833 113503835 113503974
CFEL : 113369904 113370714
CLEC : 106661079 106667579
DPX : DAPPUDRAFT_299576 DAPPUDRAFT_70042
PPOI : 119107244 119108265 119108271
CBR : CBG_23262(Cbr-pyc-1)
SHX : MS3_00005353(PYC1_1)
OVI : T265_03273 T265_03274
NVE : 5500503 5520726 5520838
VCN : VOLCADRAFT_102707(pyc1)
MPP : MICPUCDRAFT_70187(PCB)
SCE : YBR218C(PYC2) YGL062W(PYC1)
SEUB : DI49_1244 DI49_1907
SPAO : SPAR_B03160 SPAR_G01790
VPO : Kpol_461p4 Kpol_534p47
CGR : CAGL0F06941g CAGL0K06787g
NCS : NCAS_0D03180(NCAS0D03180) NCAS_0H01470(NCAS0H01470)
NDI : NDAI_0F01780(NDAI0F01780) NDAI_0I01710(NDAI0I01710)
TPF : TPHA_0B02650(TPHA0B02650) TPHA_0J01790(TPHA0J01790)
TBL : TBLA_0A09990(TBLA0A09990) TBLA_0F02480(TBLA0F02480)
TDL : TDEL_0G03550(TDEL0G03550)
KAF : KAFR_0A04660(KAFR0A04660) KAFR_0G02260(KAFR0G02260)
KNG : KNAG_0F02530(KNAG0F02530) KNAG_0M00810(KNAG0M00810)
SPAA : SPAPADRAFT_131550(PYC1)
CAL : CAALFM_C403940CA(PYC2)
NTE : NEUTE1DRAFT150551(NEUTE1DRAFT_150551)
TRE : TRIREDRAFT_21957(pyc1)
SAPO : SAPIO_CDS2380 SAPIO_CDS3574
ANG : ANI_1_440184(An04g02090)
URE : UREG_02831 UREG_04749
ZTR : MYCGRDRAFT_73616(PYC)
TVS : TRAVEDRAFT_22061 TRAVEDRAFT_72201
LBC : LACBIDRAFT_185474(PYC)
ABP : AGABI1DRAFT112162(AGABI1DRAFT_112162) AGABI1DRAFT78556(AGABI1DRAFT_78556)
ABV : AGABI2DRAFT193409(AGABI2DRAFT_193409) AGABI2DRAFT208578(AGABI2DRAFT_208578)
MRR : Moror_5660 Moror_6913
SCM : SCHCO_02494203(SCHCODRAFT_02494203)
MLR : MELLADRAFT_43264 MELLADRAFT_46050
PTI : PHATRDRAFT_30519(PYC1) PHATRDRAFT_49339(PYC2)
FCY : FRACYDRAFT_183259(PYC1) FRACYDRAFT_186955(PYC2)
TPS : THAPSDRAFT_11075 THAPSDRAFT_269908
AAF : AURANDRAFT_20291(PYC) AURANDRAFT_65502
EHX : EMIHUDRAFT_558314(PYC1)
PAEV : N297_5621(oadA) N297_5622(accC)
PAEI : N296_5621(oadA) N296_5622(accC)
PAU : PA14_71720(oadA) PA14_71740(accC)
PAP : PSPA7_6223(oadA) PSPA7_6224(accC1)
PAG : PLES_58301 PLES_58311
PAF : PAM18_5555 PAM18_5556
PNC : NCGM2_6210(oadA) NCGM2_6211(accC)
PAEB : NCGM1900_6288(oadA) NCGM1900_6289(accC)
PDK : PADK2_28910 PADK2_28915
PSG : G655_28580 G655_28585
PRP : M062_28630 M062_28635
PAEP : PA1S_29355 PA1S_29360
PAER : PA1R_gp3369 PA1R_gp3370
PAEM : U769_29860 U769_29865
PAEL : T223_29820 T223_29825
PAES : SCV20265_6166 SCV20265_6167
PAEU : BN889_06037 BN889_06038
PAEG : AI22_04625 AI22_04630
PAEC : M802_5619(oadA) M802_5620(accC)
PAEO : M801_5486(oadA) M801_5487(accC)
PRE : PCA10_55950(pycB) PCA10_55960(pycA)
PPSE : BN5_4309(oadA) BN5_4310(pycA)
PFUW : KF707C_56370 KF707C_56380
PALC : A0T30_01435 A0T30_01440
PCQ : PcP3B5_02750(cfiB) PcP3B5_02760(cfiA)
PMUI : G4G71_03430 G4G71_03435(oadA)
PPAA : B7D75_28200(oadA) B7D75_28205(accC)
PPU : PP_5346(pycB) PP_5347(pycA)
PPG : PputGB1_5395 PputGB1_5396
PPW : PputW619_5124 PputW619_5125
PPB : PPUBIRD1_0016(oadA) PPUBIRD1_0017(accC)
PPUH : B479_26465 B479_26470
PPUT : L483_31865 L483_31870
PPUN : PP4_54070(pycB) PP4_54080(pycA)
PPUD : DW66_5766 DW66_5767
PFV : Psefu_4412 Psefu_4413
PMON : X969_25685 X969_25690
PMOT : X970_25320 X970_25325
PMOS : O165_022885 O165_022890
PPJ : RK21_04984 RK21_04985
POR : APT59_21935 APT59_21940
PMOL : CLJ08_13185 CLJ08_13190(oadA)
PST : PSPTO_5510(oadA) PSPTO_5511
PSYR : N018_25555 N018_25560
PSP : PSPPH_5142(oadA) PSPPH_5143
PSAV : PSA3335_00790 PSA3335_00795
PAMG : BKM19_029270(oadA) BKM19_029275
PCI : PCH70_51180 PCH70_51190
PAVL : BKM03_29505(oadA) BKM03_29510
PVD : CFBP1590__5377(pycB) CFBP1590__5378(cfiB)
PCAB : JGS08_26340(oadA) JGS08_26345
PCOF : POR16_24910(oadA) POR16_24915
PTRE : I9H09_00550 I9H09_00555(oadA)
PSYI : MME58_25890(oadA) MME58_25895
PAST : N015_01130 N015_01135(oadA)
PLIJ : KQP88_24660(oadA) KQP88_24665
PFL : PFL_6157(oadA) PFL_6158(accC_1)
PPRC : PFLCHA0_c61180(pycB) PFLCHA0_c61190(cfiB)
PFE : PSF113_5849(oadA) PSF113_5850(accC2)
PFN : HZ99_17255 HZ99_17260
PFO : Pfl01_5639(oadA) Pfl01_5640(pycA)
PFS : PFLU_6070(oadA) PFLU_6071(accC)
PFC : PflA506_5374(oadA) PflA506_5375(accC_1)
PPZ : H045_20710 H045_20715
PMAN : OU5_3519(accC) OU5_3520
PTV : AA957_14765 AA957_14770
PCG : AXG94_15060 AXG94_15065
PVR : PverR02_31070 PverR02_31075
PAZO : AYR47_06515 AYR47_06520
POI : BOP93_26030 BOP93_26035
PRX : HRH33_00570(oadA) HRH33_00575
PMED : E3Z27_28090(oadA) E3Z27_28095
PFW : PF1751_v1c54830(pycB) PF1751_v1c54840(pycA)
PFF : PFLUOLIPICF708580 PFLUOLIPICF708585
PFX : A7318_27035 A7318_27040
PEN : PSEEN5495(oadA) PSEEN5496(accC-2)
PLUL : FOB45_25475 FOB45_25480(oadA)
PBA : PSEBR_a5592 PSEBR_a5593
PBC : CD58_29505 CD58_29510
PPUU : PputUW4_05383(pycB) PputUW4_05384(pycA)
PDR : H681_24670 H681_24675
PSV : PVLB_25325 PVLB_25330
PSK : U771_31395 U771_31400
PKC : PKB_0169(cfiB) PKB_0170(pycB)
PCH : EY04_29890 EY04_29895
PCZ : PCL1606_00610 PCL1606_00620
PCP : JM49_00570 JM49_00575
PFZ : AV641_22540 AV641_22545
PLQ : AA042_10655 AA042_10660
PALK : PSAKL28_52110 PSAKL28_52120
PRH : LT40_16545 LT40_16550
PSW : LK03_05755 LK03_05760
PPV : NJ69_19350 NJ69_19355
PSES : PSCI_1832 PSCI_1833
PSEM : TO66_31255 TO66_31260
PSEC : CCOS191_5240(pycB) CCOS191_5241(cfiB)
PPSY : AOC04_19990 AOC04_19995
PSOS : POS17_6118 POS17_6119
PKR : AYO71_00390 AYO71_00395
PFK : PFAS1_11430 PFAS1_11435
PANR : A7J50_5805 A7J50_5806
PPSL : BJP27_14410 BJP27_14415
PSET : THL1_5686 THL1_5687
PSIL : PMA3_29800 PMA3_29805
PYM : AK972_6003 AK972_6004
PADE : C3B55_00018(cfiB) C3B55_00019(cfiA)
PKE : DLD99_28630(oadA) DLD99_28635
PALL : UYA_23950 UYA_23955
PUM : HGP31_21190(oadA) HGP31_21195
POJ : PtoMrB4_54910 PtoMrB4_54920
PGG : FX982_01559 FX982_01560
PPSH : G5J76_02720(oadA) G5J76_02725
PGY : AWU82_06750 AWU82_06755(oadA)
PLAL : FXN65_27490(oadA) FXN65_27495
PSEJ : HNQ25_02200 HNQ25_02205(oadA)
PVK : EPZ47_29585(oadA) EPZ47_29590
PEZ : HWQ56_28500(oadA) HWQ56_28505
PBZ : GN234_17795(oadA) GN234_17800
PKH : JLK41_24585(oadA) JLK41_24590
PTY : JWV26_09400(oadA) JWV26_09405
PMAO : PMYSY11_4431(pycB) PMYSY11_4432(accC)
PATA : JWU58_00925 JWU58_00930(oadA)
PDW : BV82_2238(oadA) BV82_2239
PZE : HU754_000655(oadA) HU754_000660
PMOE : HV782_000670(oadA) HV782_000675
PCAM : HNE05_18425 HNE05_18430(oadA)
PSEP : C4K39_3507 C4K39_3508
PVW : HU752_000660(oadA) HU752_000665
PTRT : HU722_0000545(oadA) HU722_0000550
PXA : KSS93_02250 KSS93_02255(oadA)
PSAM : HU731_015770(oadA) HU731_015775
PTZ : HU718_000655(oadA) HU718_000660
PQI : KH389_26855(oadA) KH389_26860
PSHH : HU773_000670(oadA) HU773_000675
PXN : HU772_024640(oadA) HU772_024645
PHV : HU739_018045 HU739_018050(oadA)
PPRG : HU725_022590(oadA) HU725_022595
PNN : KEM63_16695 KEM63_16700(oadA)
PANH : HU763_024315(oadA) HU763_024320
PASG : KSS96_00565(oadA) KSS96_00570
PFAK : KSS94_26820(oadA) KSS94_26825
PMUY : KSS95_03345 KSS95_03350(oadA)
PMAM : KSS90_25190(oadA) KSS90_25195
PAZE : KSS91_00390(oadA) KSS91_00395
PALV : KSS97_00610(oadA) KSS97_00615
PPHA : BVH74_04895 BVH74_04900
PORY : EJA05_27835(oadA) EJA05_27840
PWZ : J7655_00255 J7655_00260(oadA)
PTK : EXN22_01240(oadA) EXN22_01245
PTW : TUM18999_60670 TUM18999_60680
PZA : HU749_000570(oadA) HU749_000575
PIE : HU724_000660(oadA) HU724_000665
PWY : HU734_001130 HU734_001135(oadA)
PTAE : NCTC10697_04448(cfiA) NCTC10697_04449(cfiB)
PGF : J0G10_00610(oadA) J0G10_00615
PCAV : D3880_22235(oadA) D3880_22240
PJU : L1P09_24970(oadA) L1P09_24975
PHF : NLY38_24730(oadA) NLY38_24735
PPHN : HU825_06125 HU825_06130(oadA)
PIZ : LAB08_R59830(oadA) LAB08_R59840
PPEG : KUA23_29710(oadA) KUA23_29715
PSII : NF676_27200(oadA) NF676_27205
PSIH : LOY51_26635(oadA) LOY51_26640
PCAS : LOY40_28915(oadA) LOY40_28920
PFIT : KJY40_00600(oadA) KJY40_00605
PSA : PST_0186(accC-2) PST_0187(oadA)
PSZ : PSTAB_0250(accC) PSTAB_0251(oadA)
PSR : PSTAA_0248(accC-2) PSTAA_0249(oadA)
PSC : A458_20350 A458_20355
PSJ : PSJM300_01275 PSJM300_01280
PSH : Psest_4084 Psest_4085
PSTU : UIB01_20860 UIB01_20865
PSTT : CH92_01090 CH92_01095
PCHL : LLJ08_19505(oadA) LLJ08_19510
PBM : CL52_19780 CL52_19785
PKG : LW136_21370(oadA) LW136_21375
SDEG : GOM96_11460 GOM96_11465(oadA)
AVN : Avin_02050(pycA) Avin_02060(pycB)
AVL : AvCA_02050(pycA) AvCA_02060(pycB)
AVD : AvCA6_02050(pycA) AvCA6_02060(pycB)
ACX : Achr_2030(pycA) Achr_2040(pycB)
EMO : DM558_15145 DM558_15150(oadA)
EAZ : JHT90_03320 JHT90_03325(oadA)
HPEG : EAO82_02230(oadA) EAO82_02235
HAES : LO767_16340 LO767_16345(oadA)
HARR : HV822_08760(oadA) HV822_08765
PBAU : OS670_11325(oadA) OS670_11330
MHC : MARHY2617(accC) MARHY2618(oadA)
MAD : HP15_3934 HP15_3935(pycB)
MBS : MRBBS_2714(cfiB) MRBBS_2715(pycB)
MPQ : ABA45_12830 ABA45_12835
MARI : ACP86_10415 ACP86_10420
MARA : D0851_13690 D0851_13695(oadA)
MARJ : MARI_03700(cfiA) MARI_03710(cfiB)
SDK : KHX94_11945 KHX94_11950(oadA)
SYK : KDN34_14685(oadA) KDN34_14690
SAVI : JYB87_02955 JYB87_02960(oadA)
HMI : soil367_08995(oadA) soil367_09000
ALKM : NKI27_15570 NKI27_15575(oadA)
CATT : OLW01_14985 OLW01_14990(oadA)
SPOI : IMCC21906_01785 IMCC21906_01786
SNAN : I6N98_11250(oadA) I6N98_11255
STAW : NCG89_02745(oadA) NCG89_02750
ZAL : AZF00_11690 AZF00_11695
LPO : LPO_0532(dcoA) LPO_2943(accC)
LPU : LPE509_00366 LPE509_02757
LPF : lpl0507(dcoA) lpl2591
LPP : lpp0531(dcoA) lpp2718
LPC : LPC_0475(accC) LPC_2878(dcoA)
LPA : lpa_00712 lpa_03896(accC)
LFA : LFA_0408(cfiB) LFA_2657(pycB)
LHA : LHA_0572(pycB) LHA_2961(accC)
LOK : Loa_00519(accC) Loa_00778(pycB)
LCD : clem_01600(cfiB) clem_12125(cfiA)
LLG : 44548918_00443(cfiA) 44548918_02476(accC_2)
LIB : E4T55_05540(oadA) E4T55_09700
LJR : NCTC11533_00441(cfiA) NCTC11533_02631(accC_2)
LWA : SAMEA4504053_0768(cfiA) SAMEA4504053_3168(accC_2)
LSS : NCTC12082_01813(accC_1) NCTC12082_02647(cfiA)
LANT : TUM19329_05070 TUM19329_31280
TMC : LMI_0266(accC) LMI_0806(pycB)
MDN : JT25_007875 JT25_007880
MDH : AYM39_19220 AYM39_19225
MKO : MKLM6_3889 MKLM6_3890
METL : U737_02020(oadA) U737_02025
MAH : MEALZ_2339(accC) MEALZ_2340(oadA)
MBUR : EQU24_05845(oadA) EQU24_05850
MEIN : methR_P1353 methR_P1354
MISZ : MishRS11D_08090 MishRS11D_08100
MEJ : Q7A_1446(accC) Q7A_1447(oadA)
MPIN : LGT42_011300(oadA) LGT42_011305
NWR : E3U44_14635 E3U44_14640(oadA)
ALV : Alvin_1668 Alvin_1669
ATEP : Atep_17960 Atep_17970
TVI : Thivi_0050 Thivi_0052
TMB : Thimo_1967 Thimo_1968
MPUR : MARPU_11480 MARPU_11485
TSY : THSYN_19970 THSYN_19975
TTP : E6P07_06335(oadA) E6P07_06340
CJAP : GWK36_06805 GWK36_06810(oadA)
THIM : KFB96_23175 KFB96_23185(oadA)
TBOG : LT988_05440(oadA) LT988_05445
HHC : M911_09600 M911_09605
EBS : ECTOBSL9_2867 ECTOBSL9_2868
TNI : TVNIR_1370(oadA_[H]) TVNIR_1371
TTI : THITH_07660 THITH_07665
APRS : BI364_08465 BI364_08470
GAI : IMCC3135_13170(cfiB)
TTC : FOKN1_1828 FOKN1_1830
THIC : TspCOW1_08220 TspCOW1_08240(oadA)
HCH : HCH_06704(oadA2) HCH_06705(accC2)
HAHE : ENC22_29375(oadA) ENC22_29380
HALO : BWR19_07735 BWR19_07740
HAF : C8233_06890(oadA) C8233_06895
CMAI : BFX80_12960 BFX80_12965
CAMH : LCW13_04305 LCW13_04310(oadA)
COBE : CLAM6_08380 CLAM6_08390
MMW : Mmwyl1_3925 Mmwyl1_3926
MARD : IBG28_02980 IBG28_02985(oadA)
MFOI : JSY38_12265 JSY38_12270(oadA)
MPRF : J8N69_06110(oadA) J8N69_06115
MRHI : KDW99_17820(oadA) KDW99_17825
TOR : R615_14925 R615_14930
TWAN : HUF19_02050 HUF19_02055(oadA)
OAI : OLEAN_C15090 OLEAN_C15100
MARS : A8C75_02420 A8C75_02425
MSEC : LN244_10650(oadA) LN244_10655
MRZ : KDW95_13875 KDW95_13880(oadA)
BSAN : CHH28_06670(accC) CHH28_06675(oadA)
BMAR : HF888_01990 HF888_01995(oadA)
VCW : GJQ55_11710(oadA) GJQ55_11715
OME : OLMES_5310 OLMES_5311
ACII : C4901_08795(oadA) C4901_08800
ATY : A9R16_009005 A9R16_009010(oadA)
PSPI : PS2015_1024 PS2015_1025
REV : HUE57_11865(oadA) HUE57_11870
ENM : EBS_1371 EBS_1372(pycA)
RPJ : N234_29240 N234_30500
CNC : CNE_1c12350(pyc1) CNE_BB1p09460(pyc1) CNE_BB1p09790(pyc2)
CBW : RR42_s0887 RR42_s0997
PLG : NCTC10937_05220(cfiA) NCTC10937_05221(cfiB)
LMIR : NCTC12852_01160(cfiB)