Betta splendens (Siamese fighting fish): 114849209
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Entry
114849209 CDS
T08231
Symbol
lama2
Name
(RefSeq) laminin subunit alpha-2 isoform X1
KO
K05637
laminin, alpha 1/2
Organism
bspl
Betta splendens (Siamese fighting fish)
Pathway
bspl04510
Focal adhesion
bspl04512
ECM-receptor interaction
bspl04820
Cytoskeleton in muscle cells
Brite
KEGG Orthology (KO) [BR:
bspl00001
]
09130 Environmental Information Processing
09133 Signaling molecules and interaction
04512 ECM-receptor interaction
114849209 (lama2)
09140 Cellular Processes
09144 Cellular community - eukaryotes
04510 Focal adhesion
114849209 (lama2)
09142 Cell motility
04820 Cytoskeleton in muscle cells
114849209 (lama2)
BRITE hierarchy
SSDB
Ortholog
Paralog
Gene cluster
GFIT
Motif
Pfam:
Laminin_G_1
Laminin_EGF
Laminin_G_2
Laminin_B
Laminin_N
Laminin_I
Laminin_II
Laminin_G_3
DUF948
Zw10_N
Motif
Other DBs
NCBI-GeneID:
114849209
NCBI-ProteinID:
XP_028996222
EnsemblRapid:
ENSBSLG00000010917
UniProt:
A0A6P7LQF6
LinkDB
All DBs
Position
24:11451278..11549871
Genome browser
AA seq
3259 aa
AA seq
DB search
MERLALALLAACLLRSCGAQQRGLFPAVLNLASMADITANATCGSMGPEMFCKLVEHVPG
QPVTNPQCRICNQKSQIPLEHHPIEYAIDGTNRWWQSPSIQNGMDYHYVTITLDLKQVFQ
IAYVIVKAANSPRPGNWILERSVDGVTFDPWQYYAITDTECLTRFNINPRKGPPSYTRDD
EVICTSFYSKIHPLENGEIHASLINGRPSAEDPSPTLLNFTSARYIRLVFQRIRTLNADL
MTLTFRDPKEIDPIVTRRYYYSIKDISIGGMCICYGHAKACPLNTVTKKFSCECEHNTCG
ESCDRCCPGYNQQPWMAGTFLTRHVCEECNCHGKADECYFNQTVADLSLSLDTRGKRRGG
GVCIGCRENTAGINCESCVAGHYRPTEVSAEEENPCVPCECDPQGSISQSCVADSSQATS
TQPAGSCRCKEGFGGLQCDRCAVGYMGYPSCQRCNCSVEGSSNDDPCVTPCVCKENVEGD
NCDRCKLGFYNLQGDNRRGCEKCSCMGIASQCSASAWAYQNETTLTGWHLIGDTGEKVWT
VHRHTPPYLSVRHADVVNGLGSVYYWNAPERYLGNKLSAYGGSVVFTVSYSTDQPDEMAI
RVSSEPDLIIEGGGIRIIDRRFGQPVYPSSPSTKHVALLPENFLVAETAQPITRRDFLSV
LAKVDRVMVRASYSTEFSAVYRLHSFSMEVANPSARGEKTASAVEMCSCPAGYAGTSCEA
CISGFRRVNGNLYNGVCEACRCHGHATRCHQVTGHCLDCSHHTTGLYCETCLFGYYDNAT
RGSPADCQPCACPLTLSSNSFSPTCHLSEGGELLCNQCRPGYAGPRCNRCSNGYHGNPTV
PGGSCQPCDCHGNLDLSIPGSCDPTTGQCLRCRQGYGGASCDSCAEGYYGDAITAKNCQP
CQCHTNGSVSEVCSQETGQCQCRENVVGRQCDECMPNCWWDAERQECMPCRCSRHGSISQ
RCDVEGRCICRPGYVGRRCDLRRQSYERRETRRPVERVPLEAAHARLGGLTRSGGCPRGA
YRPRTGPQTHGITTGGACVPCHCNSFGSKSFDCDETGQCRCQPGVSGPKCDRCSQGFFNF
QEGGCTPCQCSHVGNNCDANTGQCICPPNTIGERCDRCAPNHWGHDITTGCKECGCSVIG
SVTQQCNVNTGCCFCRDSFRGEKCNECQIGYRDFPQCTQCECNLPGSDTQTCDMERKVCA
CADRTGKCSCKVNVAGHNCDQCKPDTFGLSVRNPLGCSKCYCYGLTHSCTEAQGLIRMWL
TLKPEQTVLTLVDKSNTVETRRGVSFQHPEILANADLVTESVSEPYYWKLPEQFKGSMIT
AYGGQLKYAVYYEARDEVGPSSYEPQVIIKGGPNHNILITRHITGLQIGQLTRHEIDMTE
HLWKYADGRSMTREDFMDVLFYVDYILIKAAHGNLMRHSRISEIILTVAEEGRPTRESEK
AHQIEKCDCPVGYSGFSCEECAAGFYRLRAGSLAPAPASRVPTAAGMGSCVQCQCSGHSS
TCDPETSICQKCQDNTEGDRCERCAPGFYGVVRGSPDDCKPCACPLTNPKNNFSPTCVAE
GFDDYRCTACPEGYEGKYCERCATGYHGNPRTPGGHCEECKCSSWGALPGPCDPVTGQCS
CRAGALGRSCEQCMDRHVCGPTGIISCDDECSGLLISDMDRLYRIITEVTLTTPLPPPYK
LLYRFENMTEELKHMLSPQRAPERLLQLADSNLGSLVVEMDQLHSRATKVSADGEQVEDD
ADRIHKRADDLEQFIRDTLLGAKDLQLKAEELNRTLSRGDGTPEKSLKEMKEDIQAMLAE
MRRRQLGGKKGIAEEEMALAEELYQKVKRMFGDPHQATEDLKSEIKGKLSDHEGKLREAE
DLLHSAQGKTRQAASLTEQNQANLTALERKHSAVSGVKQEAQKVLGEAEDLLDEANQLSD
NINKEIEDLQEMERELDPLHDQLDDRVSHLTSGLSNGRLANHVLNAKEHAKQLNESAAIL
DGILAEAKNLSFNATAAFKAYTNIKANIDAAEKEAKAARQTANEALALALGPEVPVKEAA
RVALQKSHSLLNEANQLHKDVKANADNVAGLKGRVKAARDKTKDLLKAVNGTIATLHTIP
NDTAAKLAATKAVAADANATALDVLERLRDLNLRIHGLKRNYSELEDNVNAANQMIQDPE
KNIHAAGAKVKDLEDEADRLLEKLQPIKQLQDNLRRNISQIKELINQARKQANSIKVSVS
SGGDCLRSYRPDIRKGRYNTIILHVKTTTPDNLLFYLGSAKYVDFLALEMRKGKVSFLWD
VGSGVGRVEYPHHTIHDGNWHRIEASRNGLNGTISVYPLEGPMAGMMPTPASSTSPAAFT
ILDVDQNAYLFVGGIIGTVKRADAVRATTFTGCMGETFLDGKPIGLWNYRERQGDCKGCV
VSPQRSEGEGTVQLDGEGYAAVGRPTRWNPNVSTITFKFRTFSSDALLMYLATADMKDFM
SLELSEGKVKVNFDLGSGVGRAISASRHNDGHWKSLTMSRNKKQATVTVVDVDSSIEEKI
VATSQGSATGLNLKENQKIYFGGLPTIGNYSMAARSEVILKRYAGCLRDIEVSRTPYNLL
SSSDYTGVTKGCNVENLYTVSFSKPGYMELGGLSLGVDTEISLSFSTLADTGTILLAVGG
ASTHSHTGRNANPAGSKRRRRQSGEPHLSVMLNKGSLEVLVFTGSHNPRRVTRRPEQGAL
NDGREHSLRIERLAGRSFTVQVDEEAKREAPLPNDQPIGLQRIFLGGIPAEVEQATNRAN
VPFQGCIWNLMINTVLSDFSRPVSFDNAEIGQCPNLAPPPPAPAPTDEKEPEEEEMEPKP
VGPTPSPAPFKPHREPSAEATPPAPPVVTPTAAAAPPTAKQEASADLDSCASAVAPTALE
AAFQFGLTRNSHMSFNFNDTKVRERLILEFDLRTKEESGLVLYMARINHADFVSIQIKDG
QVCLGYDLGHGNISGCVPYSINDGNWHKIRVNRNKQRGLLTVDGRYTKQMTSPKTADLLD
VVGVIYVGGLPQNYATKRIGPILYSINGCIRNFKMVGAPVSSKASGRSDAAWEDFQFNMA
TPTARHMVGRCFVSTETGTYFEGTGFLKAVSPYRVGLDVSMSFEFRTSRTSGVLLTISNQ
VSDGLGLEIAEGKLLFHVDNGAGRITAEHAPDGEGFCDGQWHSVVARKIRHKVELVVDEK
QSQAESPNPRSNTCDTNDPIYVGGFPDGVRQVALSTNTPFQGCMRNLKIMKAAKTMDVQF
NKALELKGVQPLSCPAAAA
NT seq
9780 nt
NT seq
+upstream
nt +downstream
nt
atggagcgactcgcgctggcgctgctggccgcgtgcctgctgcggagctgcggggcgcag
cagagaggactgttcccggcagtgctgaacctggcatcaatggcagacataacagctaac
gctacctgtggttcaatgggtccagaaatgttctgtaaactggtggaacacgtaccaggc
cagcccgttacaaaccctcaatgtcggatatgcaaccaaaaaagccaaataccacttgag
catcatcccatagaatatgccattgacgggaccaaccgctggtggcaaagtccgtctata
caaaatggaatggactaccactatgtgactatcacgctggacttgaaacaggtgttccag
attgcttacgtgattgtcaaggcagccaactcacctcgcccaggtaactggatcttggag
cgctcagtggatggcgttacctttgacccctggcagtactacgctatcaccgacacagaa
tgcctgacacgattcaacattaatccacgcaaaggacctccatcatacacaagagacgat
gaagttatctgcacatctttttattcaaagatccaccccttggagaatggagagatccac
gcgtctttgatcaatgggcgtcccagtgcagaggacccctccccgaccctgctgaacttc
acctccgcccgctacatcaggctggtgtttcagcgaatcagaacgctgaacgctgacctc
atgactctgacatttcgagaccccaaagaaattgatccaattgtgacaagacggtactac
tattcaattaaggacatttccattggagggatgtgtatctgctacggccatgccaaagcc
tgtccactcaacactgttacaaagaagtttagttgtgaatgtgaacacaacacatgtggg
gagagttgtgatcgctgttgccctggttacaaccagcagccctggatggcaggaaccttc
ctcactcgacatgtctgtgaagagtgtaattgccacggtaaggcagacgagtgctacttc
aatcagaccgtagcagacttatcgctcagcctggacacccgtggcaaacgcagggggggt
ggagtttgtattggatgccgtgaaaatacggctggcattaactgtgagagctgcgtggct
ggacactacagacctactgaggtaagcgctgaggaggagaacccttgcgtcccctgtgaa
tgtgacccacaaggttctatcagccagtcatgtgttgccgattcaagccaggctacatcc
acccagccagcagggtcatgccggtgtaaggagggttttgggggtctgcaatgtgacagg
tgcgctgtgggttatatgggctacccgtcctgtcaacgctgtaactgcagtgtggaaggg
agcagcaacgatgacccatgtgtgacaccctgcgtgtgcaaggaaaacgttgagggagac
aactgtgaccgctgtaaactggggttctacaatctccagggtgacaatcgacgtggttgt
gagaagtgctcctgcatgggcatcgctagtcagtgttctgcctccgcttgggcctatcaa
aacgagaccacactaacaggctggcacctgatcggtgacacgggagagaaagtgtggact
gttcacaggcacacgcctccgtacctgagtgtccgacacgcagacgttgtaaatggcctc
ggctctgtctactactggaacgctcctgagcgatacctggggaacaagctttcagcatat
ggagggagtgtggtcttcactgtgtcctattccacagaccagccagatgaaatggccatt
agagtcagctcagaacccgacctgatcatagaaggtggtgggatcaggattatagacaga
agatttggccagcccgtgtacccatcatcaccaagcaccaagcacgttgctctgctgcca
gagaacttcctagttgcagagacagcccagccgataactcgaagggatttcctgtctgtg
ttagccaaagtggacagagtgatggtccgggcctcttacagcacagagttcagtgctgta
tacagacttcactcgttctccatggaagtagctaatccctccgccagaggcgagaagact
gcatcagctgtagagatgtgttcttgtcctgcaggatatgctggaacttcctgtgaggca
tgtatttcaggctttcggagggttaatggtaatctgtacaacggcgtatgtgaagcctgt
cgttgccatggacacgcgacgcggtgccaccaggtcacgggacactgcctggactgctca
caccacaccacaggcctctactgtgaaacctgcctttttggttattatgacaacgccacc
cgtgggtctcccgctgactgccagccttgtgcctgcccacttactctctctagcaacagc
ttcagccccacctgccacctgagtgaaggaggagagttgctgtgcaaccagtgtcggcct
ggatacgcaggaccacgatgtaacagatgttccaatggataccatggcaatcccactgta
ccaggaggctcctgccagccctgtgattgccatggaaacctggacctatccataccaggc
agctgtgatccaaccaccggccagtgtctgcgctgtcgccaaggttatggcggtgcatcg
tgtgatagctgtgctgagggttactacggcgacgccatcacagcaaaaaactgccagccc
tgtcagtgccatactaatggatctgtatctgaagtctgcagtcaggagactggacagtgc
cagtgcagagaaaatgtggttggacgacagtgtgacgagtgcatgcctaattgctggtgg
gatgctgagcgtcaggagtgtatgccctgccgctgcagccgtcacggctccatatcccaa
cgttgtgatgtcgagggccgctgcatctgtcgccccggctacgtgggccgacgctgcgac
ctgcgtcgccagagttacgagaggcgggagacgcggcggccggtggagcgggttcccttg
gaagcggcgcacgcgagattgggagggttgactcgaagcgggggatgtccccggggggcg
tacaggcccaggacggggcctcagactcacggcatcactacaggtggagcgtgtgtcccg
tgccactgcaactcattcggctccaagtcgtttgattgcgatgagaccggtcagtgccgc
tgtcagccgggcgtgagcgggcccaaatgtgaccgatgctcacagggattcttcaacttt
caagagggaggctgcacaccttgtcagtgcagtcatgtgggcaataactgtgatgccaac
acgggacagtgcatctgtcccccaaacactattggtgagaggtgtgaccgctgcgctcca
aaccactggggccatgacatcacgactgggtgcaaggaatgtggttgcagcgtgatcggc
tcagtgacccagcagtgtaacgtcaacacaggctgctgcttctgtcgtgactccttcaga
ggagagaaatgcaatgaatgtcaaattggatacagggacttcccacagtgcacacagtgc
gagtgtaacctaccaggatcggacacccagacctgtgacatggagagaaaagtgtgtgcc
tgcgctgatcgaactgggaaatgtagctgcaaggtgaatgtggcaggacacaactgtgac
cagtgcaagccagatacatttgggttgtctgttcgaaatccgctgggctgcagcaaatgt
tactgttacggcctgacacactcctgcacagaggcacaagggctcatcaggatgtggttg
acgctgaagccagagcagactgtgctcaccctggtggataagtccaacacggtggagacc
aggagaggagtgagcttccagcacccagagatcctcgctaatgcagacctggtcaccgaa
agcgtctcggagccttactactggaaactgcctgagcagttcaaagggtctatgatcacc
gcatacggtgggcagttgaaatatgctgtgtattacgaggccagagatgaagttggtcct
tcctcctatgagcctcaggttatcattaagggcggtcctaaccacaacatcctgatcaca
cgccacataacaggactccagattggtcagctcacgcgccatgagatcgacatgacagag
catttgtggaaatatgcagatggcaggtctatgactcgagaggacttcatggacgttttg
ttctatgtggactacatcctaattaaggcagcacacggcaacctaatgagacacagccgt
atttcagagatcattctcacagtggctgaggaaggacgtccgacccgagagagtgagaaa
gctcatcagatcgagaagtgtgactgtccagtcggatactccggcttttcttgtgaggag
tgtgctgcagggttctaccgcctccgtgccggctccctggcaccggccccggcgtccaga
gtgcccaccgctgccggcatgggcagctgcgttcagtgccagtgcagtgggcactcgtcc
acctgcgatccagagacgtccatctgccagaaatgtcaggacaacacagaaggcgatcgc
tgtgaacgctgtgctcctgggttctatggagtggtgcgaggttcccctgacgactgcaaa
ccctgcgcctgccccctcaccaacccaaagaacaatttcagtcctacctgtgtggccgag
ggatttgatgactaccgatgcacagcctgtcctgagggatatgagggcaaatattgtgag
aggtgtgccaccggttaccatggtaacccacggacgccaggaggccactgcgaggagtgt
aagtgttcctcgtggggggcgttgccaggaccgtgtgaccctgttaccggccaatgcagc
tgcagggccggggcgcttgggaggtcgtgcgagcagtgcatggacaggcatgtgtgtgga
cccactgggatcatctcttgtgatgacgagtgctcgggcctgttgatcagcgacatggac
cgtctgtatcgcatcatcactgaagtcactctgaccacccccctcccgccaccttacaag
ctgctgtaccgctttgagaacatgaccgaggaactaaagcacatgttgtcgcctcagaga
gctccagagagattactgcagctggccgactccaacctcggatcacttgtcgttgagatg
gaccagctacacagcagggccaccaaggtgtctgctgatggagagcaggttgaagatgat
gccgacaggattcataagcgagccgatgacctggagcagttcatcagggacacgctgctg
ggagctaaagacctccagctgaaggctgaggagctgaaccggaccctctcccggggagac
ggaaccccggagaagagcctgaaagagatgaaggaggacatccaggccatgctagcagag
atgaggaggcgacagctaggagggaagaagggcatcgcggaggaggagatggccctcgca
gaggagttgtatcagaaggtcaagaggatgttcggcgatcctcatcaggccacagaagac
ctgaaatccgagataaaaggcaagctctccgaccatgaagggaaactccgggaggccgag
gacctgcttcactctgcacagggaaagacgagacaggctgcttcactcacagaacaaaac
caggccaacctcacagccctggagaggaagcactctgcagtcagtggggtgaaacaggaa
gcacagaaagtccttggagaggcagaggatctactggatgaagccaaccagctttcagac
aacatcaacaaagagatagaggacttgcaggagatggaaagagagctggatcctcttcac
gaccagctggacgatagagtcagtcacctcaccagtggtctgagtaacggcagactggct
aatcacgtgctcaacgcaaaagaacatgctaagcagctgaacgagtctgctgccatcctg
gatggtattttggctgaagctaaaaatctgtctttcaatgcaacggctgccttcaaggcc
tacacgaacataaaggccaacattgatgcagcagagaaagaagcaaaggcagcaagacag
acggcaaatgaggctctggcgctggctttaggcccagaagtcccagtaaaggaagcagct
cgagttgcactgcaaaaaagccacagtttgttgaatgaagctaatcaactgcacaaggat
gtcaaagcgaatgcagataatgtggccgggctcaagggcagagttaaagctgcgagagac
aaaaccaaagacctgctcaaagctgtcaatggaaccatagcaacgctccacaccatcccc
aacgacactgcagctaagctagcagccaccaaggctgtggcagcagacgctaacgccaca
gcgttggacgtcctcgagcgcctcagggatctgaacctgcggatccacggcctgaagagg
aactacagcgagctggaggataatgtgaatgcagccaatcagatgatccaggacccagag
aaaaacattcatgctgcaggagctaaggtgaaagacctggaggatgaagccgacaggctg
ctggagaagctgcagccaatcaagcagctgcaggacaacttgaggaggaacatctcgcaa
atcaaggaactgatcaaccaagccaggaagcaagcgaactccatcaaagtgtctgtgtcg
tcgggtggagactgtctgcgtagttaccgccccgatatcaggaaaggaaggtacaacacc
atcatccttcatgttaaaaccaccacacctgataatctgctgttttacctgggatcagcc
aaatatgtggatttcttggccttggagatgcgtaaggggaaggtgtccttcctgtgggat
gtgggctcaggcgtgggccgcgttgaatatccccatcacaccatccacgatgggaactgg
cacagaatcgaggcgtcacgtaatggtttgaatggcactatatcagtatatcctctggaa
ggccccatggctggtatgatgcctactccagccagctccacctcacccgcagccttcacc
atcctggacgtggaccagaacgcgtacctgtttgtaggaggaataatcggaacagtcaag
agggctgatgctgtaagagcgacaacattcacaggctgcatgggggagacattcctggat
gggaagcctatcgggctgtggaactacagagagagacagggagactgtaaaggatgtgtg
gtcagcccccagcgctcagaaggcgaaggcaccgtccagctggatggtgaaggctatgct
gcagtgggcagacccaccaggtggaatcccaacgtttccaccatcaccttcaaattccgc
accttctcctccgacgctctgctcatgtacctggccaccgcagatatgaaggactttatg
tccctggagctgtcagagggcaaagtcaaggtgaactttgacctggggtcaggagttggc
agagccatatcagctagccgccacaacgatgggcactggaagtccctcaccatgtcccgg
aacaagaaacaagccacggtgaccgttgtggacgtcgacagtagtattgaggagaagatc
gtggccacgtcacaaggttctgctactgggctcaatctgaaggaaaatcagaaaatatac
tttggaggcctgcccacaattggaaactacagcatggcagccaggtcagaggtcatactg
aagcggtatgcaggttgtcttcgtgacattgaagtctcaaggactccatacaaccttctc
agcagctcagattacactggggtcactaaaggatgtaatgtagagaacctctacacagtg
agcttctctaagccggggtacatggagctgggaggcctgtctctgggggtggacacagag
atcagcctgtcgttcagcactctggcagatacagggaccatactgctggcagtgggcggc
gcttcaacacacagccacacgggccggaacgcgaaccctgccggctccaagaggagacgc
agacagtcaggagagccccatctctcagtcatgctgaacaaaggctctctggaggtcctt
gtgtttaccggcagccacaatccccgacgtgtcaccaggcggccagagcaaggcgcactg
aatgatggaagggagcactcgctgcgcatagagaggctcgctggcaggtcatttacagtt
caggttgatgaggaggcaaagagggaggcgccgcttcccaacgaccagccaatcggcttg
cagcgcatcttcctcggtggtattcctgcagaagtggagcaggcgacgaacagagccaac
gtcccattccagggatgtatatggaacctcatgatcaacaccgttctctcagatttctcc
cggcctgtgtcctttgacaacgctgagattggccagtgccccaaccttgctccacccccg
cccgcacctgctccaacagacgaaaaggagcccgaggaggaagagatggagcccaaaccc
gtcggccccactccgagtcccgctccgttcaaaccccatcgagagccttctgctgaggcc
acgccccctgctccacctgttgtcacgcctacagctgctgctgccccgcccactgctaaa
caggaagccagcgccgacttggactcctgtgcatcagctgtggctccgacggcgctggag
gcggcctttcagttcggactgaccagaaacagccacatgagcttcaacttcaatgacacc
aaagtccgagagaggctcatactggagttcgatctgagaaccaaggaggagtccggcctc
gtcctctacatggcgagaatcaaccacgctgactttgtttccatccagatcaaagatggg
caggtctgtctgggttatgacctcggtcatggaaacatatctggatgcgtcccttactcc
atcaacgacgggaactggcacaagatccgtgtcaatcgtaacaagcagagaggtcttctc
acggtggatgggcggtacactaaacagatgaccagtccaaagacggctgacctgctggat
gtggtgggagtgatttatgttggtggactgccgcaaaactacgccactaaaagaatagga
ccaattctctatagcatcaatgggtgcatccgtaactttaagatggtgggagctcctgtg
agcagcaaggcgtcaggccgctctgacgctgcctgggaggacttccagttcaacatggcc
acgcccactgcccgtcacatggtgggcaggtgcttcgtttccacagagacggggacctac
ttcgaaggcaccggtttcctaaaagcagtgtctccttacagagtgggtctagatgtctcc
atgtcgttcgagttccgaaccagcagaaccagtggagtacttctgaccatcagtaaccaa
gttagcgatggactgggactagagatcgcagagggcaagctgctcttccatgtcgacaac
ggagctggacgcatcacggcagagcacgcgcccgacggcgagggcttctgcgacggacag
tggcactcggtcgtcgccaggaagattcgccacaaggtggagctggtggtggacgagaag
cagagccaagcggagagtccgaacccacgctccaacacatgtgacaccaacgaccccatc
tacgttggagggttcccggatggcgtccgacaggtggcgctgtccaccaacactcccttc
caaggttgtatgaggaacctgaagatcatgaaggctgctaagaccatggacgtgcagttc
aacaaggctttggagcttaaaggagtccagcctctgtcctgtcctgctgcagctgcctaa
DBGET
integrated database retrieval system
