Nocardia cyriacigeorgica: NOCYR_1266
Help
Entry
NOCYR_1266 CDS
T01756
Name
(GenBank) Putative fatty acid synthase
KO
K11533
fatty acid synthase, bacteria type [EC:2.3.1.-]
Organism
ncy
Nocardia cyriacigeorgica
Pathway
ncy00061
Fatty acid biosynthesis
ncy00074
Mycolic acid biosynthesis
ncy01100
Metabolic pathways
ncy01212
Fatty acid metabolism
Module
ncy_M00082
Fatty acid biosynthesis, initiation
ncy_M00083
Fatty acid biosynthesis, elongation
Brite
KEGG Orthology (KO) [BR:
ncy00001
]
09100 Metabolism
09103 Lipid metabolism
00061 Fatty acid biosynthesis
NOCYR_1266
00074 Mycolic acid biosynthesis
NOCYR_1266
09180 Brite Hierarchies
09181 Protein families: metabolism
01004 Lipid biosynthesis proteins [BR:
ncy01004
]
NOCYR_1266
Lipid biosynthesis proteins [BR:
ncy01004
]
Fatty acid synthase
Bacteria type
NOCYR_1266
BRITE hierarchy
SSDB
Ortholog
Paralog
Gene cluster
GFIT
Motif
Pfam:
Fas1-AflB-like_hel
Acyl_transf_1
FAS_AT_central
MaoC_dehydratas
DNA_pol_B_N
ketoacyl-synt
Ketoacyl-synt_C
SAT
Fas_alpha_ACP
FAS_I_H
Motif
Other DBs
NCBI-ProteinID:
CCF62066
UniProt:
H6R4J7
LinkDB
All DBs
Position
1393810..1403079
Genome browser
AA seq
3089 aa
AA seq
DB search
MLDRLLNGAPYALAFGGQGAQWLGELEEIGRDAALEPELTALVNEAAALLEPVAEQLLVV
RPMGFDPIGWMLEDELVDADQGETSAAPSEQVLRSATVSMPGVMLSQLAALRALRLQGLD
PAEHAPMAIIGHSQGRLAASAVEAGGARDAELLAIAQLIGAAAGLVARRRGLMPVGQRSP
MVAVSNVDPDELRAVVAEVSEGVDPAIAAVVSIRNGRRRAVLSGTPAQLERVRLRCQQIH
AEQTREREAKVRGGAVFAPVFEDVDVDVAFHHPALADTVELVAGWAGQCGLDTELAGRLA
KEILVDPIDWVATVDAVVSEGAQWILDLGPGDLLSRVTGGSLKGTGVGIVAVATRAGQRS
LLTPGAAPEVAAPWSAYAPKPVRLPNGRIVVETAFTRLTGRSPILLAGMTPTTVDAKIVA
AAANAGHWAELAGGGQVTEQIFTDRVAELKTLLQPGRSVQFNSLFLDPYLWKLQLGGKRL
VQKARAAGAPFDGVVVTAGIPELEEAVALVEELTEAGITHVAFKPGTVAQIRAVLRIADE
IPGYPVIMHIEGGRAGGHHSWEDLDDLLLETYAELRARDNVLVCVGGGIGTPERATEYLT
GTWSQPHGYPVMPLDGVLVGTAAMATLEATTAPEVKQMLVDTPGTPDWVGAGTATGGMAS
GRSQLGADIHEIDNAASRTGRLLDEVAGDADAVAARRDEIIAALDATAKPYFGDLATMTY
IEWLERYVELAVGLDRRKDFDCGSDLGDAILEATRSVWLDITWRDRFAEMVRRTESRLNP
ADRGEIPTLFSDDDLERPVAALCELRKQYPAVAETLLHPADVPFFISLCKTPGKPVNFVP
VVDADVRRWWRSDSLWQAHDPRYSADQVCIIPGTVAVAGITRVDEPVGELLDRFEQDTAY
SLVRAGVVPAAVDARRTASVTSGPIDVVLAAPDVEWSGRTTVNPVHRLGDLAEWTIDGQG
AVHPPTGATLVETTGPVAEHNYVELTVPLYAGNVVRIRITVPASVYNGGAPVITADDADA
AMSALLAVAAGQSLPEVKGRVAHVNLAWTPDLIADHAGVTGSGLPADLSTLGRTVPDVLV
GACWPAVFAVLGATRTDAAESVIEGMLDLVHLDHQIELVRELPGVTSILVVRAESTSVLD
TDMGRVVEVKVTVGEMRDQGLEVVPLAELTERFAIRGRNGSGELTDPPRAAGTLSEQATD
TPRRRRRDVTIVAPRTMAAFAAVSGDHNPIHTSEAAAKLAGLGSPIVHGMWLSAAAQHAV
SAVDPQSSVPARTLTAWTTRFLGMVRPGAEIDVRVERVAVDQGSEVVEVSCRTGGDLVMT
ATGRTAAPKTVYAFPGQGIQRKGMGLDARSRSKAAKEIWDRADKHTRAALGFSILAVVRD
NPTYLKARGVEHRHPDGVLHLTQFTQVAMATLGVAQVAELREAGAFVEGAYLAGHSVGEY
NALAAVAGVLPLEAVLEVVFQRGSAMHELVPRDAKGRSNYRMAAIRPSQFGLPDDEVIGF
VSGLSEQTGEFLEVVNLNLRGSQYAIAGTVAGLEHLEAEIDRRRAEFGGKRAFILVPGID
VPFHSTVLREGVPEFRHKLEQLLPANLHPEVLVGRYIPNLVPKPFSLEREFIAEIADLVP
SEPLAAVLADFDSWSQRPTELCRVVLIELLAWQFASPVRWIETQDLLFTDTAHGGLGVER
FVEIGLGATPTVANLASQTLKLPAFGTTTVEVLNIEREAGIVYATDTDPAPVDEPDEAPA
ETAAAAPAPAAAPAPAAAPAASGGPRPDDIAFTAADATRVLIALWTKLRIDQIGPVDTIE
GLCDGVSSRRNQLLVDLGAELSLGAIDGAADADMGALSATVDRLARTYKPFGSVLTDAIG
DHLRKVFGPSGKRPAAIAERVKKVWELGDGWASHVTAEVSLGTREGASVRGGDLGGLVSG
ALSDAASVDAAIDAAVQAVAARRGISVALPATGGGGGATVDAAALGEFTEQITGRDGVLA
SAARLVLEQLGLSEEVSAPEATEDGLVDLVSAELGSDWPRLVAPAFDARKAVLIDDRWAT
AREDLARLWLADDAETAKQSVDGFTGAGEAVAAQAKWWRQRAMHEARSVLAGHYERIATA
ALSTEEPGLWSADVAVITGASKGSIAAAITGRLLGGGATVVVTTSGLDDNRLGFYRELYR
ANARNGAALWVVPANMASYTDIDALIEWVGTEQVQTAGGAKIKVKDALTPTLLFPFAAPR
VAGDLSDAGARAEMEMRVLLWSVERLIGGLSKLGADHDVDAKLHVVLPGSPNRGLFGGDG
AYGEAKAALDAVVAKWRAEKSWSSRVTLVHALIGWVRGTGLMGHNDPMVEAVEQAGVQTW
STGEMADELLKWATARARAVTATAPQQIDLTGGLAGAKLDLPALAKQAMEAEQAAAQDAD
AEQSQLIPALPAPPTLTSALPVPEWGPVTADLSDMVVIVGAGELGPYGSARTRFEMEVSD
ELSAAGVLELAWTTGMVTWENEPKPGWYDTETGEYVPESELAGKYHDAVVERCGIRRYED
DGAMIDNTSPLMTSVFLDQDLSFTVGSEAEARAFHAADPEHTVITPVAESGDWTVTRKAG
TEIRVPRKAKLSRTVGGQIPTGWDPTVWGISADMASSVDRVALWNIACTVDAFVGSGFSP
AELMSWVHPSLVANTQGTGMGGMSSMRSLYVDNLLGEPRPNDILQEALPNVALAHVVQSY
VGSYGAMVHPVAACATAAVSVEEGVDKIKLGKAEVVVAGGFDDLGIEGIVGFGDMSATAD
SAAMSAKGISDRYFSRANDRRRGGFVESQGGGTVLLARGDIAVELGLPVLGVVAYAQSFA
DGVHTSIPAPGLGALGAGRGGRESRFAAELRKLGVGADDIAVISKHDTSTAANDPNESEL
HERLAEAIGRSDGAPLFVISQKSLTGHAKGGAAAFQLIGLCQVLENGVVPPNRSLDCVDE
KMREYPHLVWVREALRFGERFPLKAGLVTSLGFGHVSGLLAVVHPEAFIQAIDPDRREDY
QRRAQQRQLAGRQRFVEAMCGGAPLYERPADRRLGGDGTPAAQVRGLEAGVLLSGEARLG
ADGVYRIGGRGCETGQLVDGSGTAGRATP
NT seq
9270 nt
NT seq
+upstream
nt +downstream
nt
ttgctggatcgtctgctgaacggggcgccgtacgcgctggcgttcggcggtcagggggcc
cagtggctgggcgagctggaggagatcggccgcgacgcggcactggaaccggagctcacc
gcgctggtgaacgaggccgcggcgctgctcgagcccgttgccgagcagttgctggtggtg
cggccgatgggcttcgacccgatcggctggatgctggaagacgagctggtcgacgccgat
cagggcgagacctccgcggcgccgtcggagcaggtgctgcgctcggccaccgtgtcgatg
cccggtgtgatgctgagccagcttgccgcactgcgcgcgctgcggttgcagggcctcgac
ccggccgagcacgcgccgatggcgatcatcggccattcgcagggccggctggccgcctcg
gcggtggaagcgggcggcgcgcgtgatgccgaactgctcgccatcgcccaattgatcggc
gccgcagcgggtctggtggctcgccggcgcgggctgatgccggtggggcagcgttcgccg
atggtggcggtgtccaatgtcgatcccgacgagctgcgcgcggttgtggccgaggtgtcc
gagggcgtcgatccggcgatcgccgctgtggtctcgattcgcaacgggcgcaggcgcgcc
gtgctgtcgggcacgcccgcgcagctggagcgggtgcggctgcggtgccagcagatccat
gccgagcagactcgtgagcgcgaggcgaaggtgcgcggcggtgcggtgttcgcgcccgtc
ttcgaggacgtcgacgtcgacgtcgccttccaccacccggcgctggccgacaccgtcgag
ctggtcgccggctgggccggccaatgcgggctggacaccgagctggcgggccggctggcc
aaggagatcctggtcgatccgatcgactgggtcgccaccgtcgacgccgtggtgtccgag
ggcgcgcaatggattctggacctgggcccgggcgatctgctgtcgcgggtgaccggtggc
tcgctgaagggcaccggcgtcggcatcgtcgcggtcgccacccgcgcgggccagcgcagc
ctgctgacgccgggcgccgcgcccgaggtggcggcgccgtggtcggcgtacgcgccgaag
ccggtgcggctgcccaacggccgcatcgtggtcgagaccgccttcacccggctgaccggc
cgctcgccgatcctgctcgcgggcatgaccccgaccacggtggacgccaagatcgtcgcg
gccgccgccaatgccggccactgggccgagctggccggcggtggccaggtcaccgagcag
atcttcaccgaccgcgtcgcggaactgaagacgctgttgcagccgggccgttcggtgcag
ttcaactcgctgttcctggatccgtacctgtggaagttgcagctgggcggaaagcggttg
gtgcagaaggctcgcgcggcgggcgctccgttcgacggtgtcgtggtcaccgccggtatc
ccggagctcgaggaagccgtcgcgctggtcgaggaactgaccgaggccgggatcacccat
gtggcgttcaagccgggcaccgtcgcgcagatccgcgcggtgctgcggatcgccgacgag
atccccggctacccggtgatcatgcacatcgagggcggccgcgccggtgggcaccactcc
tgggaagacctcgacgatctgctgctcgagacctacgccgagctgcgcgcccgcgacaat
gtcttagtctgcgtcggcggcggtatcggcaccccggagcgggccaccgaatacctgacc
ggtacctggtcgcagccgcacggctacccggtgatgccgctggacggcgtgctggtcggc
accgccgccatggcgaccctggaggccaccacggcccccgaggtcaagcagatgctggtc
gacacccccggcaccccggactgggtcggcgcgggcaccgccaccggcggcatggcctcg
ggccgcagccagctcggcgccgatatccacgagatcgacaacgccgcctcgcgcaccggc
cgcctgctcgacgaggtcgccggtgacgccgacgccgtggccgcgcgccgggacgagatc
atcgccgcgctcgacgccaccgccaagccgtacttcggcgatctggccaccatgacctac
atcgaatggctggaacgctacgtcgaactcgccgtcggcctggaccggcgcaaggacttc
gactgcggcagcgacctcggcgacgccatcctggaagccacccgttcggtgtggctggac
atcacctggcgtgaccgtttcgcggaaatggtgcgccgcaccgaatcccggctcaacccc
gccgaccgcggcgagatcccgacgctgttctccgacgacgatctcgagcgtccggtcgcg
gcgctgtgcgagctgcgcaagcagtacccggccgtcgccgagaccctgctgcacccggcg
gacgtgccgttcttcatctcactgtgcaagaccccgggcaagccggtgaacttcgtcccg
gtggtcgacgccgatgtgcggcgctggtggcgctcggattcgctgtggcaggcccacgat
ccgcgctactcggccgatcaggtgtgcatcatccccggcaccgttgccgtcgcgggcatc
acccgggtcgacgagccggtgggtgagctgctcgaccgcttcgaacaggacaccgcgtac
tcgctggtgcgcgccggtgtggtcccggcggcggtggacgcgcgccgcacggcgtcggtc
acctcgggtccgatcgatgtggtgctggccgcgccggatgtcgaatggtccgggcgcacc
accgtcaacccggtgcaccggctcggcgatctggccgagtggaccatcgacggccagggt
gcggtgcatccgccgaccggcgccacgctggtcgagaccaccggcccggtggccgagcac
aactacgtcgagctcacggttccgctgtacgcgggcaatgtggtccggatccggatcacc
gtgcctgcctcggtgtacaacggcggcgcgccggtgatcaccgccgatgacgccgacgct
gccatgtccgcgctgctcgcggtcgccgcggggcagtcgctgcccgaggtcaagggccgt
gtcgcgcacgtcaacctggcctggacccccgatctgatcgccgaccacgccggcgtcacc
gggtcgggcctgcccgccgacctcagcacgctcggccgcaccgtgcccgatgtgctggtc
ggcgcctgctggcccgccgtcttcgcggtgctcggcgcaacccgcaccgacgccgccgaa
tcggtgatcgagggcatgctcgatttggtgcacctcgatcaccagatcgagctggtgcgc
gaattgcccggcgtcaccagcattctggtggtgcgcgcggaatcgacctcggtgctcgac
accgatatgggccgcgtcgtggaggtcaaggtcaccgtcggcgagatgcgcgatcagggc
ctcgaggtggtgccgctggccgagctcaccgaacgcttcgccatccgcggccgcaacggc
tccggcgagctgaccgatccgccgcgcgcggcgggcacgctgtccgagcaggccaccgac
accccgcgtcgtcgtcgccgcgacgtcaccatcgtcgcgccgcgcaccatggccgcgttc
gccgccgtctccggcgatcacaaccccatccacaccagcgaggccgccgccaagctggcc
gggctcggcagcccgatcgtgcacggtatgtggctgtcggccgccgcccagcacgcggtg
tcggctgttgatccgcagagttcggtgcccgcccgcacgctgacggcctggaccacccga
ttcctcggcatggttcgccccggcgcggagatcgacgtgcgcgtcgagcgcgtcgcggtg
gaccagggcagcgaggtcgtcgaggtgtcctgccgcaccggtggcgatctggtgatgacc
gccaccggccgcaccgccgcaccgaagaccgtctacgccttcccgggccagggcatccag
cgcaagggcatgggcctggatgcgcgctcgcggtccaaggcggccaaggagatctgggat
cgtgccgacaagcacacccgcgccgcgctcggcttctccattctcgccgtggtccgggac
aacccgacctacctgaaggcgcgcggcgtcgagcaccggcacccggacggcgtgctgcac
ctgacccagttcacccaggtcgccatggccactctcggtgtggcgcaggtggccgaactg
cgcgaggccggcgcgttcgtcgaaggcgcctacctggccgggcactcggtcggtgagtac
aacgcgctcgccgcggtggccggagtgctgccgctggaggcggtgctcgaggtggtgttc
cagcgcggttcggcgatgcacgagctggtgccgcgtgatgccaagggccgcagcaactac
cggatggccgcgatccggccctcgcagttcgggctgccggacgacgaagtcatcggattc
gtctccggcctgtcggagcagaccggggagttcctcgaggtcgtcaacctgaacctgcgc
ggctcgcagtacgcgatcgcgggcacggtggccgggctcgagcacctggaggccgagatc
gaccggcgccgcgccgaattcggtggcaagcgcgcgttcatcctggtgcccggcatcgac
gtgccgttccactcgaccgtgctgcgtgagggtgtgccggagttccggcacaagctcgag
cagctgctgccggcgaacctgcacccggaggtcctggtcgggcgctacatcccgaacctg
gtgcccaagccgttctcgctggagcgcgagttcatcgccgagatcgccgatctggtgccg
tcggagccgctggccgcggtgctggccgatttcgactcctggtcgcagcggcccaccgaa
ctgtgccgggtggtactgatcgaactgctggcctggcagttcgccagcccggtgcgctgg
atcgagacccaggacctgctgttcaccgataccgcgcacggcgggctcggcgtggagcgg
ttcgtggagatcggtctcggcgcgacgccgacggtggccaacctcgccagccagaccctg
aagctgccggcgttcggcaccaccaccgtcgaggtgctcaatatcgagcgcgaggcgggc
atcgtgtacgccaccgataccgacccggcgccggtcgacgagcccgacgaggctcccgcc
gaaaccgccgccgccgctccggctcccgccgcggccccggctccggctgccgcaccggcc
gcctcgggtggtccgcgtccggacgacatcgccttcaccgccgccgacgccacccgcgtg
ctcatcgcgctgtggaccaagctgcgcatcgaccagatcggcccggtcgacaccatcgaa
ggactgtgcgacggcgtctcctcccgccgtaaccagctgctggtcgacctcggtgcggag
ctctcgctcggtgccatcgacggtgccgcggacgccgatatgggcgctctgtcggccacc
gtcgaccggctggcccgcacctacaagccgttcggctccgtgctgaccgacgccatcggc
gatcacctgcgcaaggtgttcggcccctccggcaagcggcccgccgcgatcgccgagcgg
gtcaagaaggtgtgggaactcggtgacggctgggccagccacgtcaccgccgaggtctcg
ctcggcacccgcgagggcgccagcgtgcgcggcggtgatctgggcgggctcgtctccggc
gccctgtccgacgcggcctcggtcgatgccgccatcgacgccgccgtgcaggccgtcgcc
gcgcgtcgcggaatctcggtggcgctgcccgccaccggcggtggtggcggtgcgacggtc
gacgccgccgcgctcggcgaattcaccgagcagatcaccggccgcgacggcgtgctggcc
tcggccgcgcgcctggtgctcgagcagctgggcctcagcgaggaggtctccgcgccggag
gccaccgaggacggcctggtggatctggtctcggccgaactgggttcggactggccgcgc
ctggtcgcgcccgccttcgatgcccgcaaggcggtactgatcgacgaccgctgggccacc
gcgcgcgaggatctggcccggctgtggctggccgacgacgccgagaccgccaagcagtcg
gtggacgggttcaccggtgccggcgaggctgttgccgcccaggcgaagtggtggcggcag
cgcgccatgcacgaggcgcgttcggtgctggccggccactacgagcggatcgccacggcc
gcgttgagcaccgaggagccgggactgtggtcggccgacgtcgccgtgatcaccggtgcc
agtaagggttccatcgccgccgcgatcaccgggcgcctgctcggtggcggcgcgacggtc
gtggtcaccacctccggcctcgacgacaaccggctcggcttctaccgcgagctctaccgc
gccaacgcccgcaacggtgccgccctgtgggtggtgcccgcgaacatggcctcctacacc
gatatcgacgcgctgatcgaatgggtcggcaccgagcaggtgcagaccgcgggtggcgcc
aagatcaaggtcaaggacgcgctcaccccgacgttgctgttcccgttcgcggcgccgcgc
gtcgcgggcgatctctccgacgccggtgcccgcgccgaaatggagatgcgggtactgctg
tggtcggtggagcggttgatcggcgggctgtccaagctcggcgccgaccacgacgtcgac
gcgaaactgcatgtggtgctgccgggttcgcccaaccgcggcctgttcggtggtgacggc
gcctacggtgaggccaaggccgccctcgacgccgtggtcgccaagtggcgggccgagaag
tcgtggtcgagccgggtcaccctggtgcacgcgctgatcggctgggtccgcggcaccggc
ctgatgggccacaacgatccgatggtcgaggcggtcgagcaggcgggcgtgcagacctgg
tccaccggggagatggccgatgaactgctcaaatgggccaccgcacgggcccgggcggtc
accgcgaccgcaccgcagcagatcgacctgaccggcggcctggccggcgccaagctcgac
ctgcccgcactggccaagcaggccatggaggccgagcaggccgccgcgcaggacgcggat
gccgaacagtcgcagctgattcccgccctgccggccccgcccacgctgacctccgcgctg
ccggtgcccgaatggggcccggtgaccgccgatctcagcgatatggtcgtcatcgtcggc
gcgggcgagctcggaccctacggttcggcgcgcacccgcttcgagatggaggtctccgac
gagctgtcggcggccggcgtgctggaactggcctggaccactggcatggtgacctgggag
aacgagcccaagcccggctggtacgacaccgaaaccggtgagtacgtgcccgaatccgag
ctggccggcaagtaccacgacgccgtggtcgagcgctgcggtatccgccgctacgaagac
gacggcgccatgatcgacaacaccagcccgctgatgacctcggtgttcctcgatcaggac
ctgtcgttcaccgtcggcagcgaggccgaggcccgcgccttccacgccgccgaccccgag
cacacggtgatcaccccggtcgccgagtccggcgactggaccgtgactcgcaaggccggc
accgagatccgggtgccgcgcaaggcgaagctttcgcgcaccgtcggtggccagatcccc
accggctgggatccgacggtttggggcatctccgccgatatggcgtcctcggtggaccgg
gtggcgctgtggaacatcgcctgcaccgtcgatgcgttcgtcggctccgggttcagcccg
gccgagctgatgagctgggtgcacccgagcctggtggccaacacccagggcaccggcatg
ggcggcatgtcctcgatgcgctcgctctacgtcgacaacctgctcggcgagccgcgcccg
aacgacatcctgcaggaagcgctgccgaacgtggcgctggcgcatgtggtgcagtcctac
gtcggcagctacggcgcgatggtgcacccggtggcggcgtgcgcgaccgcggcggtgtcg
gtggaagaaggtgtcgacaagatcaagctcggcaaggccgaggtggtcgtggccggcggc
ttcgacgatctgggtatcgagggcatcgtcggtttcggcgacatgtcggccaccgcggat
tcggcggccatgagcgccaagggcatcagcgaccgctacttctcccgcgccaacgaccgc
cgccgcggcgggttcgtggaatcgcagggtggtggcacggtgctgctggcccgcggtgac
atcgcggtcgagctcggcctgccggtgctcggtgtggtggcctacgcgcagtccttcgcc
gacggcgtgcacacctccatcccggcgcccggactcggcgcgctgggtgcgggccgcggc
gggcgggaatcccggttcgccgccgaactgcgcaagctcggcgtgggcgccgacgacatc
gcggtgatctccaagcacgacacctccaccgcggccaacgacccgaacgaatcggaactg
cacgaacgcctcgccgaggcgatcgggcgctccgacggcgcgccgctgttcgtcatctcg
cagaagagcctgaccgggcacgccaagggcggtgcggccgcgttccagctgatcggcctg
tgccaggtgctggaaaacggtgtggtgccgccgaaccggagcctggactgcgtcgacgag
aagatgcgcgaatacccgcatctggtgtgggtgcgcgaggcgctgcgcttcggcgaacgg
ttcccgctcaaggcgggcctggtgacctcgctgggcttcggccacgtgtcgggtctgctc
gcggtggtgcatccggaggcgttcatccaggcgatcgacccggaccggcgcgaggactac
cagcgccgggcgcagcagcgtcagctcgccggacgtcagcggttcgtggaggccatgtgc
ggtggcgccccgctctacgagcggccggccgaccgacggctcggcggcgacggcactccc
gcggcgcaggtgcgcgggctggaggcgggcgtgctgctgtccggcgaggccaggctcggc
gcggacggggtgtaccggatcggtggacgtggttgtgagaccgggcaactcgtcgacggt
tccggcaccgcggggcgcgccacaccgtag
DBGET
integrated database retrieval system