Orcinus orca (killer whale): 101288504
Help
Entry
101288504 CDS
T05247
Symbol
DCHS1
Name
(RefSeq) protocadherin-16 isoform X2
KO
K16507
protocadherin-16/23
Organism
oor
Orcinus orca (killer whale)
Pathway
oor04392
Hippo signaling pathway - multiple species
oor04519
Cadherin signaling
Brite
KEGG Orthology (KO) [BR:
oor00001
]
09130 Environmental Information Processing
09132 Signal transduction
04392 Hippo signaling pathway - multiple species
101288504 (DCHS1)
09133 Signaling molecules and interaction
04519 Cadherin signaling
101288504 (DCHS1)
09180 Brite Hierarchies
09183 Protein families: signaling and cellular processes
04515 Cell adhesion molecules [BR:
oor04515
]
101288504 (DCHS1)
Cell adhesion molecules [BR:
oor04515
]
Cadherins
Cadherin related
101288504 (DCHS1)
BRITE hierarchy
SSDB
Ortholog
Paralog
GFIT
Motif
Pfam:
Cadherin
Cadherin_FAT4_N
Ig_CLSTN
Ig_Cad99C_N
Cadherin_CELSR2_9th
Motif
Other DBs
NCBI-GeneID:
101288504
NCBI-ProteinID:
XP_004279279
LinkDB
All DBs
Position
8:48263530..48298831
Genome browser
AA seq
3294 aa
AA seq
DB search
MQKERGIAPSCPGMQSPRPLPLLSLLLLLGAGVPGAWGQAGSLDLQIDEEQPAGTLIGDI
SAGLPAGTAAPPMYFISAQEGSGVGTDLAIDEHSGVVRTARVLDREQRDRYRFTAVTPDG
ATVEVTVRVADINDHAPAFPQARAVLQIPEHTALGTRYPLEPARDADAGRLGTQGYALSG
DGAGETFRLETRPGPDGAPVPELVVTGELDRENRSHYMLQLEAYDGGSPPRRAQALLDVT
LLDINDHAPAFNQSRYHAVVSESLAPGSPVLQVYASDADAGANGAVTYEINRRQSEGDGP
FSIDAHTGLLRLERPLDFEQRRVHELVVQARDGGAHPELGSAFVTVHVRDANDNQPSMTV
IFLSADGSPRVSEAAPPGQLVARISVSDPDDGDFAHVNVSLEGGEGHFALSTQDSVIYLV
CVARQLDREERDAYNLRVTATDSGSPPLRAEAAFVLHVTDVNDNAPAFDRQLYRPEPLPE
VALPGSFVVRVMARDPDQGTNGQVTYSLAPGAHTRWFSIDPTSGIITTAASLDYELEPQP
QLIVVATDGGLPPLISSATVSVALQDVNDNEPQFQRTFYNASLPEGTQPGTCFLQVTATD
ADSGPFGLLSYSLGAGLGASGSPPFRIDAHSGDVCTTRTLDRDQGPSSFDFTVTAVDGGG
LKSMVYVKVFVSDENDNPPQFYPREYAASLSAHSTPGTAVLRVRAHDPDQGPHGRLSYHI
LAGNSPPLFALDEHSGLLTVAWPLARRANSVVQLEIGAQDGGGLQAESSARVNVSIVPGT
PIPPVFEQLQYVFSVPEDVAPGTSVGVVQAHNPSGRLGPVTLTLSGGDPRGLFSLDGATG
LLQTLRALDRELLGPVLELEVRAGSGVPPAFAVARVRVLLDDVNDNSPAFPAPEDTVLLP
PSTAPGTPIYTLRALDPDSGVNSRVTFTLLAGGGGAFTVDPTTGHILLMGPLGPPGGPAH
ELELEARDGGSPPRTSHFRLRVVVQDLGTRGLAPHFDSPTYRVDLPSGTTPGTQVLQVQA
RAPDGGPVTYRLAADGPSSPFGLEPQTGWLWVRAALDREAQELYTLKVTAVSGSKAELGQ
QTATATVRVSILNQNDHSPRLSEEPTFLAVAENQPPGTSVGRVFATDRDSGPNGRLTYSL
RQLSEDSKAFRIHPQTGEVTTLQTLDRERQSSYQLLVQVQDGGSPPRSVTGTIHIAVLDL
NDNSPTFLQASGAAGGGLPVQVPDGVPPGTLVTTLQARDPDEGENGTILYMLTGPGSELF
SLHPHSGELLTAAPLTRAERPHYVLTLSAHDQGSPPRSSSLQLLVQVLPSARSAEPPPEP
SDPDPAAPVPVVLTVTAAEGLQPGSLLGSVAPPESAGVGALTYTLVGGADPEGTFALDAA
SGRLYLARPLDFEAGPAWRALTVRAEGPGGAGARLLRVQVRVQDENEHAPAFARDPLALA
LPENPEPGAPLYTFRASDADGPGPNSDVRYRLLRQEPPVPALRLDSRTGALSAPRGLDRE
TTPALLLLVEATDRPANASRRRAARVSARVFVTDENDNAPVFASPSRVRLSEDQPPGPAA
LHVVARDPDLGEAARVSYRLAAGGDGYFRLHPSTGALSVVRSLDREQRAEFLLTVVASDH
GSPPRSATQLLTVSVADVNDEAPTFQQQEYSVLLRENSPPGTSLLTLLATDPDLGANGQV
TYGGISGESFSLDPDTGVLTTLRALDREEQEEINLTVYARDGGSPPLLTHVTVRVAVEDE
NDHAPTFGSAHLSLEVPEGQDPQTLTTLRASDPDVGANGQLQYRILDGDPSEAFVLDLAS
GEFGTMRPLDREVEPAFQLQIEARDGGQPALSATLFVTVTVLDANDHAPAFPVPAYSVEV
PEDAPAGTLLLQLQAHDPDAGPNGRVTYYLGAGAAGAFLLEPSSGELRTATALDREQCPS
YAFTVNAVDGAAAGPLSTTVPITIMVRDVNDHAPTFPTSPLRLRLPRPGPSLSTPTLALA
TLRAEDRDAGANASILYRLAGTPPPGTTVDSYTGEIRVARSPVSLGPQDRVLFIVATDLG
RPARSATGVVIVGLQGESERGPRFPRASSEAMLRENAPPGTPIVSPKAIHTGGSSGPITY
SILSGNEKGTFSIQPSTGAVTVRSAEGLDFEANPRLRLVLQAESGGAFAFSVLTLTLQDA
NDNAPRFLRPHYVAFLPESRPLEGPLLQVEADDLDQGPSGQISYSLAASQPARGLFRVDS
ATGTITTTAILDREIWAETRLVLMATDRGSPALVGSATLTVMVIDTNDNRPTIPQPWELR
VSEDALLGSEIAQVTGNDVDSGPVLWYVLSPSGPQDPFSVGRYGGRLSLTGPLDFEQRDR
YHLQLLAHDGPHEGRANLTVLVEDVNDNAPAFSQTLYQVMLLEHTPPGSAILFVSATDRD
SGANGHVSYHLASPAEGFSVDPNNGTLFTTVGTMALGHEGPGVVDVVLEARDHGTPSRAA
RTTVHVQLQDQNDHTPSFTLSHYRVAVTEDLPPGSTLLTLEATDADGSRTHAAVDYSIVS
GNRGRVFQLEPRLAETGEGDGLGPRALGCLVLLEPLDFESLTQYNLTVAAADRGQPPRSS
AVPVTVTVLDVNDNPPVFTRASYRVAVPEDTPVGAELLHVEASDADPGPHGLVRFTLSSG
DPFGLFQLDESSGDLRLAHPLDCETQARHQLVVRAADPAGAHFALAPVTIEVQDVNDHRP
AFPLGLLSTSLAENQPPGTLVTTLHATDGDAGAFGRLRYDLLDAGPGPEGREAFALNSST
GELRARVAFDYEHTGSFQLLVGATDAGNLSASVTVSVLVTGEDEYDPVFLAPAFHFQVPE
GARRGHSLGHVQATDEDGGADGLVLYSLATSSPYFGINQTTGALYLQVDSRAPGSGTGTS
GGGGRTRREAPRELRLEVVARGPLPGSRSATVPVTVDITHTALGLAPDLNLLLVGAVAAS
LGVVVVLALAALVLGLVRARSRKAEAAPGPMSQAAPLASGSLQKLGREPPSPPPSEHLYH
QTLPSYGGPGAGGPYPRGGSLDPSHSSGRGSAEAAEDDEIRMINEFPRVASVASSLAARG
PDSGIQQDADGLSDTSCEPPAPDTWYKGRKAGLLLPGAGATLYREEGPPAAATAFLGGCG
LSPAPTGDYGFPADGKPCVAGALTAIVAGEEELRGSYNWDYLLSWCPQFQPLASVFTEIA
RLKDEARPCPPAPRIDPPPLITAVAHPGAKSVPPKPASTATTRAIFPPASHRSPISHEGS
LSSAAMSPSFSPSLSPLAARSPVVSPFGVAQGPSASALSAESGLEPPDDTELHI
NT seq
9885 nt
NT seq
+upstream
nt +downstream
nt
atgcagaaggagcggggcattgcaccttcttgcccaggcatgcagagccccaggccccta
cccctgctatcactgctgctgctcctgggcgcgggggtgccgggcgcctggggccaggct
gggagcttggacctgcagattgacgaggagcagccagcgggcacgctgatcggggacatc
agtgcagggcttccagcgggcacggcagcacctcccatgtacttcatctccgcccaggag
ggcagcggtgtgggcacagacctggccattgacgagcacagcggggtggtccgtacagcc
cgggtcttggaccgcgagcagcgggatcgctaccgcttcactgcagtcactcctgatggc
gccaccgtggaagttaccgtgcgagtggctgacatcaatgaccacgctccagccttccca
caggctcgagctgtcctgcagatacctgagcataccgctcttggcacccgctacccactg
gagcctgctcgtgacgcagacgctggccgcctaggaacccagggctatgcgctgtctggt
gatggggctggagagaccttccggctagagacacgtcccggtccagatggggccccagtg
cctgagctggtagttaccggggagctggaccgagagaaccgctcacactacatgctgcaa
ctggaggcctacgacggtggttcaccaccccggagggcccaggccctactggacgtgacg
ttgctggacatcaatgaccatgccccagctttcaatcagagccgctaccacgctgtggtg
tccgagagcctggcccccggcagtcctgtcttgcaggtgtatgcctctgatgccgatgcc
ggtgccaatggggctgtaacttatgagatcaaccggaggcagagcgagggtgacggaccc
ttctccatcgacgcacacacggggctgctgcggctggagcggccactggacttcgagcaa
cggcgggtccatgaactggtggtacaggcacgggatggaggggctcaccctgagctgggc
tcggcctttgtgaccgtgcacgtgcgagatgccaatgacaatcagccctctatgactgtc
atcttcctgagtgcggacggctccccccgagtgtctgaggctgccccccctggccagctg
gttgctcgaatctctgtgtcagacccagacgatggtgactttgcccatgtcaatgtgtcc
ctggagggtggagagggccactttgcactcagcacccaggacagcgtcatctacctggtg
tgcgtggctcggcagctggatcgagaggaaagggatgcctataacttgcgagttactgcc
acagactcaggctcacccccgctgcgagccgaggctgcctttgtgctgcatgtcactgat
gtcaacgacaacgcccctgcctttgaccgccagctctaccgacccgagcccctgcctgag
gttgcgctgcctggcagcttcgtggtgcgcgtgatggcccgggaccctgaccagggcacc
aacggtcaggtcacctatagcctggccccaggtgcccacacccgctggttctccatcgac
cccacttcaggcatcatcaccacagctgcttcactggactatgagttggaaccacagccg
cagctaattgtggtggccacggatgggggcttgccccctctcatctcctctgccacagtg
agtgtggccttgcaggatgtgaatgataatgagccgcagttccagaggactttctacaat
gcctccttgcctgagggcacccagcccggaacctgcttcctacaggtgacagccacagat
gcagacagtggcccatttggcctcctctcctattccttgggtgccggacttggggcctcg
gggtctcccccattccgcatcgatgcccacagtggcgacgtgtgcacgacccggaccctg
gaccgcgatcaggggccctcaagcttcgacttcacagtgacagctgtggatgggggaggc
ctcaagtctatggtgtacgtgaaggtgtttgtgtcagacgagaatgacaacccacctcag
ttttatccacgggagtatgctgccagtctgagcgcccacagtacaccaggcacagctgtg
ctgagggtgcgtgcccatgaccctgaccagggaccccacgggcgactctcctaccacatc
ctggctggcaatagccccccactgtttgccttggatgagcactcagggctgttgacggta
gcctggcccttggccagacgggccaattccgtggtgcagctggagatcggggctcaggat
ggaggtggcctgcaggcagagtccagtgcccgagtcaacgtcagcattgtgcctggaact
cccataccaccagtatttgagcaactacagtatgtcttttctgtgccagaggatgtggca
ccaggcaccagtgtgggtgtagtccaagcacacaatccatcaggtcgcctggggcctgtg
accctcaccctatcaggtggggatccccgaggactcttctccctagatggggccacaggg
ctgttacaaacgcttcgagctctggaccgggagctgctgggaccagtgctagagctggag
gtacgggcaggcagtggagtgcccccggctttcgccgtagctcgggtgcgtgtgctgctg
gatgatgtgaatgacaactcccctgccttccctgcacctgaagacacggtgttgctgccg
ccaagcactgccccagggactcccatctatacactgcgggctctggaccccgactcaggc
gttaacagtcgagtcacctttaccctgcttgctgggggtggcggggccttcactgtggac
cccaccacaggccacatactgcttatgggacctctggggcctccaggggggccagcccat
gaactggagctggaagcccgggatggcggctccccaccccgcaccagccactttcgactg
cgggtggtggtacaggacttggggacccgtgggctggctccccacttcgacagccctacc
taccgtgtggacctgccctcaggcaccacccctggaactcaagtcctgcaagtgcaggct
cgagcaccagatgggggccctgtcacctaccgccttgcagcagatgggccaagtagcccc
tttggcctggagccacagactgggtggctctgggtgcgagcagcactggaccgtgaggcc
caggagttatacacgctgaaagtaacggcagtgtctgggtccaaagctgagttggggcag
cagacagccacagccaccgtgagggtcagcatcctcaaccagaatgaccacagtccccgc
ttgtctgaggagcccaccttcctggctgtggctgagaaccagcccccagggaccagtgtg
ggccgggtctttgccactgaccgagactcaggacccaatggacgtctgacctacagcctg
cgacagctgtcagaagacagcaaggccttccgcatccacccccagactggagaggttacc
acactccaaactctggaccgtgagcggcagagcagctaccagctcctggtgcaggtgcag
gatggggggagcccaccccgcagtgtcacaggcaccatacacattgcagtgctcgacctc
aatgacaacagccccaccttcctgcaggcttcaggggctgctggtgggggcctccccgta
caggtaccagatggtgtgcctccaggaacgctggtgacaactctgcaggccagggatccg
gacgagggggagaatggaacaatcctatacatgctaactggtcctggctcagagctcttc
tctctgcaccctcactcaggggagctgctcactgcagcacccctgacccgagcagagcgg
cctcactatgtgctgacgctaagtgctcacgaccaaggcagcccccctcggagctccagc
ctccagctgctggtgcaggtgcttccctcagctcgctcggctgagcctccgccggaaccc
tcagacccagacccggccgcacctgtgcctgtcgtgctgaccgtgacagcagccgagggc
ctccagcccggctctctgttgggctcggtggcgccgccagagtcggcgggagtgggcgca
ctcacctacacactggtgggcggcgcggacccggagggcactttcgcgctggacgcggcc
tccgggcgcttgtacctggcgcggcccctggacttcgaggcgggcccggcgtggcgcgcg
ctcacagtgcgcgccgaggggccgggaggcgcgggcgcgcggctgctgcgagtgcaggtg
cgcgtgcaggacgagaacgagcatgcgccggccttcgcgcgtgacccgctggctctggcg
cttccggagaacccagagccgggggcgccgctgtacactttccgcgcgtcggacgccgac
ggcccgggccccaacagcgacgtgcgctaccgtctgctgcgccaggagccgcccgtgccc
gcgcttcgcctggactcacgcaccggggcgctcagcgccccgagaggcctggaccgggag
accactcccgctctgctgctgctcgtggaagccaccgaccggcctgccaatgccagccgc
cgccgcgcagcacgcgtttccgcgcgcgtctttgtcacggatgagaatgacaacgctccg
gtctttgcgtcgccctcacgtgtgcgcctctctgaggatcagccgccggggcccgcagcg
ctgcacgtggtagcccgggacccggacctgggcgaggctgcacgcgtgtcctatcgcctg
gcagctggcggggacggctacttccggctacaccccagtaccggagcgctgtccgtggtg
cggtccctggaccgtgaacagcgagctgaattcttgctgactgtggtggcctccgaccac
ggctccccgccgcgctcagccacacagctcctgactgtcagtgttgccgacgtcaacgac
gaggcacccactttccaacagcaggagtacagcgtcctcttgcgtgagaacagcccgcct
ggcacatctctgctcactctgctggcaaccgacccagacctgggggccaatgggcaagtg
acttatggaggcatctctggcgaaagcttctccctggacccagacactggagttcttaca
actcttcgggccctggatcgggaggagcaggaggagatcaatctgacagtgtatgcccgg
gatgggggctcacctccgctgttgacccatgtcacagtgcgagtggctgtggaggatgag
aatgaccacgccccaacctttgggagtgcccatctctccctggaggtgcctgagggccag
gacccacagacccttaccacgctgcgggcctctgacccggatgtgggagccaacgggcag
ctgcagtaccgcatcctggatggggacccatcagaagcctttgttctggacctagcttca
ggggagtttggcaccatgaggccactagaccgggaggtggagccagcattccagctgcag
atagaagcccgggatggaggccagccagctctcagcgctactctgtttgtgacagtgaca
gtgctggatgctaatgaccatgcccccgccttccctgtgcctgcctactctgtggaggtg
cctgaggatgcacctgcagggaccctgctgctgcagctacaggctcatgatcctgatgct
gggcccaatggccgcgtgacctactacctgggtgcaggtgcagcaggagccttcctgctg
gagcccagctctggggaattgcgcacagccacagccctggacagagagcagtgtcccagt
tatgcctttaccgtgaatgcagtggatggtgcagctgctggacccctaagcaccacggtg
cctatcaccatcatggtgcgcgatgtcaatgaccacgcgcccaccttccccaccagtccc
ctgaggctgcgcctgccccgcccaggccctagcctcagtaccccaaccctggctctggcc
actctgcgagctgaagaccgtgacgctggtgccaatgcctctatcctataccggctggca
ggcacaccacctcctggcaccactgtggactcttacactggtgaaatccgtgtggcccgc
tcccctgtatccctgggtccccaggatcgtgtcctcttcatcgtggcaaccgaccttggc
cgtccagctcgctctgccactggtgtggtcattgttgggctgcagggagagtctgagcgt
ggaccccgctttccccgggctagtagcgaagccatgctccgtgagaatgcacccccaggg
actcccattgtctcccccaaggctatccatacagggggctcaagtggacccatcacatac
agcattctcagtgggaatgagaaagggacattctccatccagcctagtacaggagccgtc
acggttcgctcagcggaggggctggactttgaggcaaacccacggctgcgactggtgctg
caggcggagagtggaggagcctttgccttctccgtgctgaccctaaccctgcaagatgcc
aacgacaatgctccccgcttcctgcggccccactacgtggccttcttgcccgagtccagg
cccttggagggacccctgctgcaggtcgaagcagatgacctggatcaaggccccagtgga
cagatctcctacagtctggccgcatcccagccagcccggggattgttccgtgtggactca
gccacaggcactatcactaccacagctatcctggaccgtgagatctgggccgaaacacgg
ctggtgctgatggccacagacagaggaagcccagccctggtgggctcagctaccctgacg
gtgatggtcatcgacaccaatgacaatcgccccaccatcccccagccctgggagctccga
gtgtcggaagatgcactattgggctcagagattgcacaggtaacagggaatgatgtggac
tcaggaccagtactgtggtatgtgctgagcccttctgggccccaggatcccttcagtgta
ggccgctatggtggccgcctctccctcacgggccccttggatttcgagcagcgtgaccgc
taccacctgcagctcctggcacatgacgggcctcacgagggccgtgccaacctcacggtg
ctcgtggaggatgtcaatgacaatgcacccgccttctcacagaccctctaccaggtgatg
ctgcttgagcacacacccccaggcagtgccattctcttcgtctctgccactgaccgggat
tcaggtgccaatggtcatgtctcctaccacctggcttcccctgctgagggcttcagtgtt
gaccccaacaacgggaccttgttcacaacagtggggacaatggccttgggccatgaaggg
ccaggagtggtggatgtggtgctggaagcacgagatcatgggacgccgtctcgggcagca
cgcaccacagtgcacgtgcagctgcaggaccagaacgaccacaccccgagcttcacattg
tcacactaccgtgtggcggtaactgaggatctgccccctggttccaccctgctcaccctg
gaggccacagatgccgacggaagccgcacccacgccgccgtggactacagcattgtcagt
ggcaaccggggccgagtcttccagctggaaccccggctggctgagactggggagggtgat
ggactaggcccccgggccctgggctgcctggtgttgcttgagcctctagacttcgaaagc
ctaacccagtacaatctaactgtggctgcagctgaccggggtcagccacctcgaagctcg
gctgtgccggtcaccgtcactgtgctggatgtcaatgataacccacctgtcttcacccga
gcatcctaccgtgtggcagtacccgaggacacacctgttggcgccgagctgctgcacgtg
gaggcctccgacgccgacccaggcccgcatggccttgtgcgtttcaccctcagctcaggc
gacccttttgggctctttcagctggatgagagctcgggtgacttgcgactggcccacccc
ttggactgtgagacccaggctcgacatcagcttgtcgtgcgggctgccgaccctgccggg
gcacactttgctctggcaccagtgaccatcgaggtccaggatgtgaatgaccacaggcca
gccttccctctgggcttgctcagcaccagcctggcagagaaccagcctccaggcactctc
gtgactactctgcatgcgacggatggcgatgctggggctttcggaaggctccgctacgac
ctgctggacgctgggccggggcctgagggccgcgaggcatttgcgctgaacagctcaaca
ggggagttgcgggcacgagtggcctttgactatgagcacacaggaagcttccagttgctg
gtgggtgctaccgatgctgggaatttgtcagcctcagtcactgtgtcagtgctggtgact
ggtgaggatgaatatgacccagtgttcttggctccagctttccacttccaagtgccggaa
ggtgcccggcgtggccacagcctgggtcacgtgcaggccacagacgaggatggaggcgct
gatggcctggtgctctattcccttgccacctcttccccctattttggtatcaaccagact
acaggtgccctgtacctgcaggtggacagccgggcaccaggcagcggaacaggcacctct
gggggtgggggccggacccgacgtgaggcaccacgggagctgaggctggaggtggtagca
cgggggcctctgcctggttcccggagtgccacagtgcccgtaactgtggacatcacccac
actgcgctgggcctggcacctgaccttaacctgctattggtgggggctgtggcggcctcc
ctgggagtcgtggtggtgctcgcattggcagccctggtcctagggctggtgcgggcccgg
agccgcaaggctgaggcggcacctggcccgatgtcacaggcagcacccctggccagtggc
tctctacagaagctgggccgagagccacccagcccaccgccttcagagcacctgtatcac
cagactctccccagctatggtgggccaggagctggaggaccctacccccgtggtggctcc
ctggacccttcacactcaagcggccgaggctcagcggaggctgcagaggatgacgagatc
cgcatgatcaacgagttcccccgtgtggccagtgtggcttcctctctggctgcccgtggc
cctgactccggcatccagcaggatgcagatggactgagtgacacatcctgcgagccacct
gcccccgacacctggtataagggccgcaaggcagggctgctgctgccgggtgcaggggcc
actctgtaccgagaggagggccccccagccgctgccacagccttcttggggggctgtggc
ctaagccctgcacccactggggactatggctttccagcagatggcaagccatgtgtggca
ggagcactgacggccattgtggctggggaggaggagctccgtggcagctataactgggac
tacctgctgagctggtgccctcagttccagccgctggccagcgtcttcacagagatcgcc
cggctcaaggatgaggctcggccgtgtcccccggctccccgtattgacccaccacccctc
atcactgccgtggcccacccaggagccaagtctgtgccccccaagccagccagcacggct
acaacccgggccatcttcccaccagcctctcaccgctcccccatcagccatgaaggttcc
ctgtcctcagctgccatgtcccccagcttctcgccctcactgtctcctctggctgctcgc
tcacctgtcgtctcgccatttggggtggcccagggcccctcagcctcagcactgagtgca
gagtccggcctggagccacccgatgacacggagctgcatatctag
DBGET
integrated database retrieval system