Piliocolobus tephrosceles (Ugandan red Colobus): 111527933
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Entry
111527933 CDS
T07491
Symbol
PCNT
Name
(RefSeq) LOW QUALITY PROTEIN: pericentrin
KO
K16481
pericentrin
Organism
pteh
Piliocolobus tephrosceles (Ugandan red Colobus)
Brite
KEGG Orthology (KO) [BR:
pteh00001
]
09180 Brite Hierarchies
09182 Protein families: genetic information processing
03036 Chromosome and associated proteins [BR:
pteh03036
]
111527933 (PCNT)
Chromosome and associated proteins [BR:
pteh03036
]
Eukaryotic type
Centrosome formation proteins
Centriole proteins
111527933 (PCNT)
BRITE hierarchy
SSDB
Ortholog
Paralog
Gene cluster
GFIT
Motif
Pfam:
PACT_coil_coil
Motif
Other DBs
NCBI-GeneID:
111527933
NCBI-ProteinID:
XP_026303200
Ensembl:
ENSPTEG00000006885
LinkDB
All DBs
Position
19:complement(33050703..33169081)
Genome browser
AA seq
3312 aa
AA seq
DB search
MELEQEQRRRKVEAGRTKLAHFRQRKTKGDGSHSEKKTAKRKGSAVDAPVQEESPIAKED
RALRGGGDICKSTSCDDTPDGAGWAFAAQLENRDGEKREDLEQLQQKQVSDHPPEQRGIF
TISDHHPEQREIFTVNDPHQEQRGIFTVSDPXXXFTVSDHHPEQREIFTVSDPHQEQRGI
FTVSDPHREQRGIFTVSDHPPEQPGIFTVSDHHPEQREIITKECEQECELAIADLQSSRE
DEAGLQQSQAVHGLELEALRLSLSNMHTAQLELTQANLQKEKETALTELREMLNSRRAQE
LALLQSRQQHELELLREQHAREKEEVVLRCGQEAAELKEKLQSEMEKNAQIVKTLKEDWE
SEKDLCLENLRKELSAKHQSEMEDLQNQFQKELAEQRAELEKIFQDKNQAESALRNLESQ
HQAAMEKVREDLQSEHCRRLEDLEFKFRESEKEKQLELENLQASYEDLKAQSQEEIRRLW
SQLDSARTSRQELSELHEQLLARASHVEDLEQLKQREKTQHESELEQLRIYFEKKLRDAE
KTYQEDLTLLQQRLQGAREDALLDPVEVGLSCVGLEEKPEKGRKAHVGELSPERRKESLP
CFQAELEESRGHLLEAPESPLCIQHEGHVSDRCCAEASALAQGWHLEPSEGHSQELPWVH
LQGVQDGALEADTEVAARVLGLETEHKVKLSLLQTELKEEIELLKIENRNLYEKLQHETR
LKENLEKAKHNLVEDHQEELNNAKQKTELMKQEFQKKEMDWKVTKEELQREAEEKLTLML
LELREKAESEKQTIVNKFELREAEMRQLQDQQAAQILDLERSLTEQQGRLQQLEQELTSD
DALRCSQCGREPPTAQDGELAALHMKEDCALQLMLARSRFLEERKEITEKFSAEQDAFLQ
EAQEQHTRELQLLQERHQQRLLSVTAELEARHQAALGELTASLESKQGALLAARVAELQT
KHAADLSALETRHLSSLDSLESCYLSEIQTIREEHRRALELLRADFEEQLQKQDSLHQTI
LTQELEKLKQKHDEELQSVWDGLQTAASRELTGTVAHELQGAHQGEFGSEKKAALHEKEE
TLQLQSAQAQPFHQEEKESLSLQLQKKNHQVQQLKDQVLSLSHEIEECRSELEALQQRRE
RENQEGANLLSMLKADVDLSHSERGALLDALRRLLGLFGETLRAAITLRSRIGERVGLCL
DDAGAGLALSAAPALEETWSDVTLPELDRTLSECAEMSSVAEMSSHMRESFLMSPESVRE
CEQPIRRVFQSLSLAVDGLLEMALDSSRQLEEARQIHSRFEKEFSFKNEETAQVVRKHQE
LLECLKEESAAKAELALELHKTQGTLEGFKVETADLKEALAGKEDSEHRLVLELESLRRQ
LQQAAQEQAVLREECTRLWSQGEATATDAEAREAALRKEVEDMTREQSETRKQAEKDRSA
LLSQMKILESELEEQLSQHRGCAKQAEAVTALEQQVASLDKHLRNQRQFMDEQAAEREHE
REEFQQEIQRLEGQLRQVAKPQPWGPHDSQQAPLDGEVELLQQKLREKLDEFNELALQKE
SADRRVLRQEEEIKRLEEMNISIRKKVAELQEEVEKQKNIMKELAQDKELLKKQQMSSLL
LASTLQSTLDAGSCPEPPPGSPPEGPEVQLEVTQSALLQRESEVLDLKEQLEKMKGDLES
KNEEILHLNLKLVMQNSQTAVRLGELQEENVSSKVIYSRSSEIEELKATIENLRENQKRL
QKEKAEEIEQLHEVIEKLQHELSLMGPVVHEVSDSQAGSLQSELFCSQAGGPRGQALQGE
LEAALEAKEALSQLLADRERGHSQALEALQQRLQGAEEAAERQLAELERSAALREAEVEG
MASRIQEFEAALKAKEAMIAERNLEIDALNQRKVAHSAELEAVLLALAHIRRALEQQPLA
AGAEPPELQRLRAQCARLSRQLQVLHQRFLRCQVELDRRQACRTTAHTWVPGAHPQPCVD
GGAKAQVPGDMEASHDGAMEPVVPDPQGDLQPVLMTLEDAPLCKQEGVMSVLTICQRQLQ
SELLLVKNEMRLSLEDVGKGKEKVLEDCQLQKVDLIAQVKQLQEKLNRLLYSMTFQNVDA
ADAKSPWPLASAHLLESSWSDGSCDGEEPDTSAPIDACDASTATGGVTDVIRNRDSLIPG
ETPDSPIQEKSECQHRSLSSPTGMLGGSRHQSHAAEAGPRKSPAGMLDLSSWSSPEVLRK
DWTLEPRPHLPVTPHSGALSLCSADTSLRDSADTSLPQTQGPGLLCSPGVSAAALALQWA
ESPPADDHRVQRMAVEKDVEDFITTSLDSQETLSSPPPGLEGKTDGSKKSDGLGFGARLS
PGSGGPEAPTAGPVTPAAISGRFQPLPETMKEKEVHPQHVKALLQMVCDESHQILALSEG
LAPLTGEPHPSRKGDEDVPTVCPDWRGELLQVVQEAFEKEQEIQGVELQPRLSGSDRGGH
SSLLERLEKVISEQGDLQEKSLEQLRLPDRSSLLSEIQALRAQLRMAHLQNQEKLQHLRT
ALTSAEARGRQQEHQLRRQVELLAYKVEQEKCIAGDLQKTLSEEQEKANGVQKLLAAEQS
VVRDLKSELCESRQENERLSRSLCEVQQEVLQLRSMLSNKENELKAALQELEREQGKGRA
LQSQLEEEQLRHLQREGQSAKALEELRASLETQRAQSSRLCVALKHEQTAKDNLQKELRI
EHSRCEALLAQERSQLSELQKDLAAEKSRTLELSEALRHERLLTEQLSQRTQEACVHQDT
QAHHALLRKLKDEKSRVADLQAMLEKVQQQALRSQQQLEAEAQEHCEALRREKEVSATLK
STVDALHTQKRELRCSLEREREKPAWLQAELEQSHPRLKEQEGRKAARRSAEAGQSPAAA
ERWRKWQRDKEKLRELELQRQRDLHKIKQLQQTVRDLESKDEVPGSRLHLGCARRAAGSD
ADHLREQQRELEAMRQRLLCAAGLLTSFASQAMDRTVNDWTSSNEKAVVSLLHTLEELKS
DLSRPTSSQKKMAAELQLQFVDVLLEDNVSLTKALSTVTREKLELSRAVSKLEKMLKHHL
QKGCSPSKLERAAWKPDETAAQSSLRRPDPGRLPPAASEEAHPSNVKMEKLYLHYLRAES
FRKALIYQKKYLLLLIGGFQDSEQETLSMIAHLGVFPSKAERKTTSRPFTRFRTAVRVVI
AVLRLRFLVKKWQEVDRKGALAQGKVPRPGPRVRRPQSPPRTRESPPTRDVPSGHTRDPA
RGHGLAAAASPHVGGRATPSPNSRLERSLTASQDPEHSLTEYIHHLEVIQQRLGGALPDS
PSKKSCNQKIKQ
NT seq
9939 nt
NT seq
+upstream
nt +downstream
nt
atggaacttgagcaagagcagcggcgccgaaaggtggaggccgggaggacgaagcttgct
cacttccgacagagaaaaacaaaaggtgacggttcgcattcggagaaaaagacggcgaag
aggaagggctcggctgtcgatgcgcctgtccaggaggagagtccgatagccaaggaggac
cgggcactccgtggaggaggggacatttgcaaaagcacatcatgtgacgacacccctgat
ggggcaggatgggcctttgctgctcagctggagaaccgagatggagagaagagagaggac
ttggaacaactgcagcagaagcaagtcagtgaccaccctccagaacagcgtgggatcttc
acaatcagtgaccaccatccagaacagcgtgagatcttcacagtcaatgacccccatcaa
gaacagcgtgggatcttcacagtcagtgacccccnnnnnnnnttcacagtcagtgaccac
catccagaacagcgtgagatcttcacagtcagtgacccccatcaagaacagcgtggaatc
ttcacagtcagtgacccccatcgagaacagcgtgggatcttcacagtcagtgaccaccct
ccagaacagcctgggatcttcacagtcagtgaccaccatccggaacagcgtgagatcatc
acaaaggagtgtgaacaagaatgtgaacttgccattgctgacctgcagagcagccgtgaa
gatgaggctggcctgcagcagagtcaggctgtgcatggccttgaactggaggcgctgcgt
ctgagtctgagcaacatgcatacggcacagctggagctgacccaggccaacctccagaag
gagaaggagacggcgctgacggagctgcgggagatgctcaacagccggcgtgcccaggag
ctggccctgctgcagagcaggcagcagcacgagctggagctcctcagggagcagcacgca
cgggagaaggaggaggtggtgctcaggtgtggacaggaagcagccgagctgaaggagaaa
ttacaatcagaaatggagaaaaacgcccagatcgtaaagaccctgaaggaagactgggaa
tctgaaaaagatttatgtttagaaaatctacgcaaagaattgtctgcaaagcatcagtca
gaaatggaggatttacaaaaccagtttcagaaagaattggcagaacagagagctgagttg
gagaagatttttcaagacaaaaaccaggctgaatcggcccttaggaacctggagagccaa
catcaggcagccatggagaaggtccgtgaagacctgcagtccgagcactgccggcgttta
gaggacttggagttcaagttcagagagagcgaaaaagaaaaacagctggagttagagaat
cttcaggcatcgtatgaagacctgaaggcacagtcacaagaagagatcaggcgcttgtgg
tcccagcttgattctgccaggaccagtagacaggaattgagtgagctacatgagcaactc
ctggcgcgagcctctcacgtggaagatttagaacagctgaagcagcgagaaaaaacccag
cacgagtccgaactggagcaactaaggatttattttgaaaagaaattaagggatgccgag
aaaacttaccaggaagacctaaccctgttgcagcagaggctgcagggggcgcgtgaagat
gctcttctggaccctgtggaagttgggttgtcctgtgtgggtttagaagagaaacctgag
aaaggaagaaaagctcacgttggtgaactcagccctgagcgacgtaaggagagcctgcca
tgcttccaggcagagttagaagaaagccgcgggcacctattggaagcgccggagtctccc
ctctgcatccagcatgaggggcacgtctcagacagatgctgcgcagaggcttcagcattg
gcacagggatggcatctggagccctccgaagggcacagccaagagcttccctgggtgcat
ctccagggtgtgcaggacggggccttggaggccgacacagaggtggcagccagagtcttg
ggtctggaaactgagcataaggttaaactctcgcttcttcagacggaactcaaggaagaa
attgaactcctaaaaatagaaaacagaaatttgtacgagaagttgcagcatgaaacccgt
ctgaaggagaatttggagaaggcaaaacacaatctagttgaagaccaccaggaggaacta
aataatgctaagcaaaagactgaactgatgaaacaggaattccaaaaaaaagaaatggac
tggaaagttacgaaggaggagctacagcgggaagctgaggagaagttaacgttgatgctt
cttgaactgagagaaaaggctgaatccgagaaacagaccatcgtaaacaagtttgagctt
cgagaagctgaaatgaggcagcttcaggaccaacaggcagcccagatcctggatctggag
aggtccttgacggagcagcagggccgcctgcagcagctggaacaggagctcacttcagac
gatgccctgcgttgcagccagtgtgggcgggagccgcccacagcccaggacggggagctt
gccgcgctccacatgaaggaagactgtgccctgcagctgatgctggcccggagcaggttt
ttagaggaacgtaaagagatcaccgagaaattcagtgcagaacaagatgccttcctgcag
gaggcccaggagcagcacacccgcgagctgcagctcctccaggagagacaccagcagcgg
ctcttgtcagtgacggcagagctcgaggccagacaccaggccgcactgggcgagctgaca
gcctccttagagagcaagcagggggctctgctggccgcgcgtgtggccgaactgcagacg
aaacacgctgccgacctcagcgctctggagaccagacatctgtccagccttgattctttg
gaatcctgttacctgtctgaaattcagaccatccgtgaggagcacaggcgggccctagag
ctcttacgagcagactttgaggaacaactgcagaaacaggactctcttcaccaaacaatt
ttgactcaagagttggagaaactgaagcaaaaacatgacgaggagctgcagtctgtgtgg
gacggcctgcaaaccgcagcgagcagggagctcaccggaaccgtggctcacgagctgcag
ggagcgcaccagggtgaatttggaagtgaaaagaaagctgctttgcatgaaaaagaggag
acacttcagctccagagtgcgcaggcacagccttttcaccaagaggagaaagagtctttg
tctctgcagcttcaaaagaagaatcaccaagtccagcagctgaaagaccaggttttatcc
ttaagtcacgagatagaagagtgccgctccgagctggaggccctgcagcagaggcgggag
cgggagaaccaagaaggtgcaaacctcctctccatgctcaaggccgacgtcgacctgtcc
cacagcgaaagaggggccctcctggacgccctgcgcaggctgctgggtttgtttggagag
acactgagagcagccatcaccctgaggagccggatcggggagcgcgtggggctctgcctg
gatgacgcgggcgcaggcctggccctgtcggcagctccggcgctggaggagacgtggtct
gatgtcaccctcccggagctggacagaactttgtctgaatgtgcagagatgtcttccgtg
gctgaaatgagcagccacatgcgtgaaagctttctcatgagcccagaaagcgtgcgggaa
tgcgagcagcccatccggagggtcttccagagcctcagcctggccgtggacggcctcttg
gagatggccctggactccagcaggcagctggaagaagcacgacaaattcattctcgtttt
gaaaaagaatttagttttaaaaacgaggagacggcacaggttgtcaggaagcaccaggag
ctgctggagtgcttgaaggaggagagtgcagcaaaggcagagctggcgctggagctgcac
aagacgcagggcaccctcgagggattcaaggtggagacggccgatctgaaggaggcactg
gccgggaaggaggattctgagcaccgtctggtcctggagctggagagtctgagacggcag
ctgcagcaggcggcccaggagcaggcagtgctgagggaggagtgcacccgtctgtggagc
cagggggaggccacagccacggacgccgaggccagagaagctgctctccggaaggaagtg
gaggatatgaccagagaacagtcggagaccaggaagcaggctgagaaggaccgctcggcc
ctgctctcccagatgaagattttggagtctgagttagaggaacagctgtctcagcatcgc
gggtgtgccaagcaggcggaggccgtcaccgccctggaacagcaggtggcgtctctggac
aagcatttgcgcaaccagcggcaattcatggatgagcaggccgctgagcgggagcacgaa
cgcgaggagttccagcaagagattcagaggctggaggggcagctccgccaggtggccaag
ccacagccctggggccctcacgacagccagcaggcaccgctggatggagaggttgagttg
ttacaacaaaagttgagagaaaagttggatgaatttaatgaattggctctacagaaagag
tcagcagatagacgagtgttgaggcaggaagaagaaattaaacgtctggaggagatgaac
atcagcatcaggaagaaagtggccgagctccaggaagaagtggaaaaacagaagaacatt
atgaaagagctggcgcaggataaagagttgttaaagaaacagcagatgagtagcttactt
ctggcatccacattgcagtctacactagatgcaggcagttgccccgagcctcctccgggc
agccctcctgagggtccagaagtacagttggaggtgacacagagcgcactcctgcagcgt
gagagtgaggttttggacttgaaagaacaactagagaagatgaaaggtgacttagaaagt
aaaaatgaagaaatactacatctgaacttaaaattagtcatgcagaacagccagactgct
gtccgcctcggagagcttcaggaagaaaacgtgagctcgaaggtcatatattccagaagt
tctgagattgaagagctgaaagccacgattgaaaatctgcgagagaaccagaaacgatta
caaaaggagaaagcagaggaaattgaacaactccatgaagtcattgagaagctgcagcat
gagctgtccctcatggggcccgtggtgcacgaagtcagcgacagtcaggctggcagtctg
cagagcgagctgttctgctctcaggccgggggccctcgggggcaggccctgcagggcgag
cttgaggctgcgcttgaagccaaggaggccctgagccagctgctggctgaccgggagcgc
gggcacagccaagccctggaggccctgcagcagcgcctccagggcgcagaggaggctgcg
gagcggcagctggctgagctggagcgcagtgcagccctcagggaggctgaggtcgaaggc
atggcctcccggatccaggagttcgaagctgccctgaaagcaaaggaagcaatgattgcc
gagagaaatttggaaattgacgcattgaaccagcggaaggtggcccactctgccgagctg
gaggccgtcctgttggccttggcccacatccgccgcgcgctggagcagcagcccctggca
gctggggcggagcctcccgagctgcagcggctccgagcacagtgtgcccgcctcagccgc
cagttgcaggtgctgcaccagcggttcctgaggtgccaggtggagctggacaggcggcag
gcctgcagaaccacagctcacacatgggtgcctggggcccacccacagccttgcgtggat
ggtggcgccaaggcccaggtccccggcgacatggaggcctcccacgatggtgctatggag
ccggttgtccctgacccacagggtgatctgcagcccgtcctgatgacgttggaggacgca
cctctctgcaagcaagaaggcgtaatgtcagtgctcaccatctgccagaggcagctgcag
tcagagctgctcttggtgaaaaatgaaatgcgcctgagtctggaggacgtaggcaagggt
aaggaaaaagtgctggaagactgtcagctgcagaaggtcgatctgatcgcgcaggtgaaa
caacttcaggaaaaattgaaccgtttgctgtattccatgaccttccagaacgtggatgct
gccgacgccaaatctccgtggcccttggcctcggcacacctgttggagagcagctggagt
gatggttcctgtgacggtgaagagcctgacacatcagcccccatagatgcgtgtgatgcc
agtacagccacggggggtgtgactgatgttatcagaaatcgggattccttgataccaggt
gaaacgccagattcccccattcaggaaaagtcagaatgtcagcacaggtccctttcttca
ccaaccggcatgcttggcggctcccgccaccagagccacgctgcggaggctgggccccgg
aagagcccggccgggatgctggatctctcttcctggagctcccctgaggtcctcaggaag
gactggaccctggagccccggccccacctccccgtgacaccccactcaggagccctgagc
ctgtgcagtgccgacacatccctgcgggacagcgcggacacctcgctaccacagacccag
gggccggggctgctttgttccccgggtgtgtccgcagcagcgctggccctgcagtgggct
gagtctccgccagctgatgaccaccgtgtgcagaggatggctgtggagaaagatgtggaa
gattttatcacaacgtctcttgattctcaagaaacattaagttcacctcctcctggatta
gaagggaaaactgatggaagtaagaaaagtgacggcttgggttttggagcaagactgagc
ccggggtcaggaggccctgaggctccaactgctggtcctgtgacccctgctgccatctct
ggaaggtttcagccactgccggaaaccatgaaggagaaggaggtacatccgcagcacgtg
aaggctttgctacagatggtgtgtgacgagagccaccagatcctggcactgtcagaaggc
cttgcaccgctgactggcgagccacacccctcccggaagggagacgaggacgtgcccacc
gtgtgccctgattggagaggggagcttctgcaggttgtgcaggaagcatttgaaaaagag
caggagatacagggggttgagctgcagccccgactcagtggctcagatcggggaggtcac
agctccctgcttgaaagactggagaaggtcatcagtgagcagggagacctgcaggaaaag
tccctggagcagcttcgcctgccggaccggagcagcctgctgtccgagatccaggctctg
cgcgcccagctgcgcatggcgcacctgcagaaccaggagaagctgcagcacttgcgcacg
gccctgacgagcgcggaggcgcgtgggagacagcaggagcaccagctgcgcaggcaggtt
gaactgctggcctataaagtagagcaggagaagtgcatcgctggcgacttgcagaagacg
ctgagtgaagagcaagagaaagcaaacggcgtgcagaagctcctggcggcagagcagagt
gtggtgcgagatttgaagtccgagctctgtgagagcaggcaggagaatgaacggctgtcc
cggtccctctgcgaggtgcagcaggaggtcctccagctgagatccatgctgagcaataag
gagaacgagctgaaggctgcgcttcaggagctggagagagagcaggggaaggggcgtgcc
ctgcagagccaactggaggaggagcagctccggcacctgcagagggagggccagagtgcc
aaggccctggaggagctgcgggcgtctctggagacacagcgtgctcagagcagtcgactc
tgtgtggcgctgaaacatgagcagacggccaaggacaacctgcagaaggagctgcgcatc
gagcactcacgctgcgaggctttactggctcaggagcggagccagctctccgagctccag
aaggaccttgctgccgagaagagccgcaccctggagctgtcagaggccttgcggcacgag
cggctcctgactgagcagttgagccagaggacacaggaggcttgtgtgcaccaggacaca
caggcccatcacgctctgctgcggaagctgaaggacgagaagtcccgggtggcggacttg
caggcgatgcttgaaaaggtgcagcagcaagccctgcgttctcagcagcagcttgaagct
gaggctcaggagcactgtgaggcgctcaggagagagaaggaggtaagtgccaccctgaag
tcgacggtggacgccctgcacacccaaaaacgagagctgagatgctctctggagagagag
agggagaagccagcgtggttgcaggcagaattagagcagtcacacccacgcttgaaagag
caagaaggacgcaaggctgcgaggaggagcgcggaggccgggcagagcccagcggctgcg
gagcggtggaggaagtggcagagagacaaggagaagctgcgagaattagaactgcagcgt
cagcgtgacttgcataagatcaagcagcttcagcagacggtgagagacctggagtcgaag
gacgaggtgcctggcagccgcctccacctgggttgtgcccgcagggctgctggctcggat
gcggaccacctccgggaacagcaacgagagctggaggcgatgaggcagcggctgctctgt
gctgccgggcttctcaccagcttcgccagccaggccatggacaggacagttaatgattgg
acatcatccaatgagaaagcagtggtgtcgttactgcacacgttggaggagctgaagtct
gacttgagcaggcccacctcctcccagaaaaaaatggcagcagagctgcagctccagttt
gtggacgtcctgctggaagacaatgtttccctcacgaaagccctcagcacggtgacccgg
gagaagctggagctgagcagggccgtgtctaagcttgagaagatgctgaagcaccatctg
cagaaaggctgcagcccgagcaagttggaaagggctgcttggaagccagacgaaacggct
gcacagagttccctgaggcgcccagaccccggccggcttccaccggctgccagcgaggaa
gcacaccccagcaacgtcaagatggaaaaattgtacctacattatttgagagcagagagc
tttagaaaagctctgatttatcaaaagaagtatcttttgctgttgattggtggtttccag
gattctgaacaagaaacactctccatgattgcccatttgggagtatttccttccaaagca
gagcggaaaaccacatctcgtcctttcacacggttccgcacggccgtcagggtggttatt
gccgtattgagattacgttttttggttaagaaatggcaagaagtagatcggaaaggagct
ctggcacaaggcaaagttcctcgcccagggccccgagtacgacggccgcagtctccaccc
agaaccagagagtcccccccaacccgggatgtgccctcaggccacaccagggaccccgcc
agaggccacggactggcagcagcagcctccccacacgttgggggaagagccactccatcc
ccaaattcaagattagaaagatccctgactgcttctcaagatccagaacattccttgaca
gagtatattcaccatttagaagtgatccagcaaagactgggaggggctctaccagattct
ccttcaaagaaatcctgcaaccagaagattaaacagtga
DBGET
integrated database retrieval system
