Seriola dumerili (greater amberjack): 111239103
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Entry
111239103 CDS
T05606
Symbol
atm
Name
(RefSeq) serine-protein kinase ATM
KO
K04728
serine-protein kinase ATM [EC:
2.7.11.1
]
Organism
sdu
Seriola dumerili (greater amberjack)
Pathway
sdu03440
Homologous recombination
sdu04068
FoxO signaling pathway
sdu04110
Cell cycle
sdu04115
p53 signaling pathway
sdu04210
Apoptosis
sdu04218
Cellular senescence
Brite
KEGG Orthology (KO) [BR:
sdu00001
]
09120 Genetic Information Processing
09124 Replication and repair
03440 Homologous recombination
111239103 (atm)
09130 Environmental Information Processing
09132 Signal transduction
04068 FoxO signaling pathway
111239103 (atm)
09140 Cellular Processes
09143 Cell growth and death
04110 Cell cycle
111239103 (atm)
04210 Apoptosis
111239103 (atm)
04115 p53 signaling pathway
111239103 (atm)
04218 Cellular senescence
111239103 (atm)
09180 Brite Hierarchies
09181 Protein families: metabolism
01001 Protein kinases [BR:
sdu01001
]
111239103 (atm)
09182 Protein families: genetic information processing
03032 DNA replication proteins [BR:
sdu03032
]
111239103 (atm)
03036 Chromosome and associated proteins [BR:
sdu03036
]
111239103 (atm)
03400 DNA repair and recombination proteins [BR:
sdu03400
]
111239103 (atm)
Enzymes [BR:
sdu01000
]
2. Transferases
2.7 Transferring phosphorus-containing groups
2.7.11 Protein-serine/threonine kinases
2.7.11.1 non-specific serine/threonine protein kinase
111239103 (atm)
Protein kinases [BR:
sdu01001
]
Serine/threonine kinases: Other
PIKK family [OT]
111239103 (atm)
DNA replication proteins [BR:
sdu03032
]
Eukaryotic type
DNA Replication Termination Factors
Other termination factors
111239103 (atm)
Chromosome and associated proteins [BR:
sdu03036
]
Eukaryotic type
Meiosis-specific proteins
DSB formation-related proteins
111239103 (atm)
DNA repair and recombination proteins [BR:
sdu03400
]
Eukaryotic type
Check point factors
Other check point factors
111239103 (atm)
BRITE hierarchy
SSDB
Ortholog
Paralog
GFIT
Motif
Pfam:
PI3_PI4_kinase
FAT
TAN
FATC
HEAT_ATR
Motif
Other DBs
NCBI-GeneID:
111239103
NCBI-ProteinID:
XP_022624615
LinkDB
All DBs
Position
Unknown
AA seq
3096 aa
AA seq
DB search
MCSTMNLALDDLRDCCRGLENDKATERKKEAERFRRLIRCPEVVQELDRTSGSKAKASKQ
LTWDTVFRFLQRYIQKETESMQSSKSVTATALATRKKKMAEMCSLIKYFIRYANKRGPRL
KCSELLRHVMEVLQSSYSCSVCGEDYSSLLVKDILSVRKYWCDIAPQQWQSLLDLYCSLF
NSSSKPINHVLVSRVIHIVVQGCCMQTDGFNNTLFGFFSKALLNARQEKYLTVLEHLVSA
LNIFLRSAAMNCRMRVCHLGEELLPSILHIWVDKRPSAALKEKIVEFFNLQICVHHPKGA
KTQDTGAHAEDWTKWRSQLYSLYDALVREISHIGSRGKYVTGTRHIAVKDNLIELTADIC
RQLFTDDNETHILEVTQASLRATQMGSPTHGAASKRRRIELGWEVLRDHLQSQQSDFDII
PWLQITAVLVSKYPSMVPGHELVPLLSVLYQLLAEQRRGERGPYVLRSLREVARCQAHHP
ERTQVHRAELGRLWGRVWALALRGVSSPQTEALSFDLLASIVHGGLIDMDREFWKLFSGS
ACKLSQSAAVCLSQALLKCPVPKRLIPSSGWDYVGVTEGSGSPNLKETLIAWLLMSDQSD
EMEDSSRPHPIICRDFPYNLIAGILVSLTLKDTRAGLRFLLGAEGAKSVFFQEQVPSDAK
DNLEEIERLYLQFSFNTLPSEVRGSEESVKPSSADGQFAAVPGLRNKLEQSLLCVADNLL
NCYSPDSQSTPPECLVRCFSLLTGVLAGYVSTGVLTEEEAYHSQLLTKAKALAQDFSGFV
STVKVKMTEEGTMHTLKSVMELCTQSASRKNKDNVCPISSSLFIMILPGSLLSELAEVLK
LLLSSVSKRVSLVDEDDHADDDRDVTRTQQGDDIDLFEDGEEAHSSTNETHRQKGDNTDA
SCGPGAKSLLAEDHLGQQDLALLAILEFLCECGSVQLVHGLLFKPQEVQRKLLLLLEQID
FTKALHLNMYLVLLKKLPAEDSLSPVEFDSLLRPLADLCNLYRQDQEICVAVLLGLLPSI
RSLGKTHHMPEEMKHVQGSLLQVVSGFCFLSQTGKCISTMRAALVQCLVALLEADPCCKW
AVLDIRGSDRMEDRLVSVILPSYLADAHHQVRMLVAMSVERLFLEMSPEQPDKKKMLPLK
HQQDAFENVYLKAQEGMRLPRSSSSEDKCDEVFNRKATLLKSLSVVLCCSPVCEKQSLFA
LFQSYKENNIEEKLIKKMLCSVSRVLGYRNVKTFVSSHLYYLVAEWLAQRQSDDRYTLGS
FPYTLLDHDTVKDFYNSSYQVLIPHLVFLDDFEQVRSIGRYLERDWKKLLADCFPKIMVN
ILPYFALSGQDSQVAQQREKAHRVYDLLKDADCLGKQQIDSLIHTNLADIVVELLMTLYE
GGEAEGDRGDLKRFIGELDPVPNPPYFSSYVIKATLDYLSKCHSANHKSLVAILSKTPIS
IQRILLAVCERAAETTNSYERHRILLMYHLFVNLLLREVKEGLGGAWAFVLRDIIYTLIH
HINGRSLQCDEVSTRSLSLCCDLLTSVCQAALQFCDDALDCHLQVIVGTLTSQVTSRPVI
SQQVLSLLQFLVIENQQKLKGAISRLEPFPDLPEFRELRSVQNRLKHNTGTFTLRQEIVH
FLSVSSCDSLPLTRLEGLKELTRQLHDNKGQIRELLKECHADPSDSVLVKLVLSLLQLSK
IAANHPGGADILEAAGGCLGELGPVDFSTIALLHGRDPFYEKAVSLFTSTELQCLYIILN
CMNDALTQPRIEVRQAAAQCVKNILATQSGVDFWEQHKDNRDPMLAYLNPFRKDKSKVVA
VSAEESSEAREKLESQEFWIPQAGGHKAWLRALCTALLDSGGVKSEALLLSRPLCLVKVD
CCQRLLPLIIHSILLDDSDGSWRELLSSHIQDFFSFCSRSAQASSRSATPLNSDSDSDTA
SQGLYDKTSLRTMLAVIDYLRHQQRPLDSNSKSCGTVCNSNFWLELNYLEVAKAAQSCAA
HFTALLYTEIYVDKIKANMEESRRTKSRATYKLSFEEHSQKFTISSLTEKSMEDTNISLQ
ELLIEVYRSIGEPDSLYGCGGETMTSSLTRIRTYEHEAMWGKALTSYDLHSTLPEATRQV
GIVVGLQNFGLSSILATYMRGLESEGVEWGAELRELRFQSAWRNTQWDCELSERSDKSTP
GFHESVFCSLQALRNKEFSIFDETLKQARGAEVEELCRGSLEAVSSLYPALRNLQSIREL
ESVKQLFSRPFSDVTLRDVCSQWRQHSQLLADSDFALVEPILTVRSVAQNTLMSRVGDPD
SIQYLSSVLTDHLMELCQLARKAGNTQRAEQAVFRMKQHGGGGSWASSPVLSWQLEEAQV
FWAKGEQGLALGLLRQMIHNLEKRVDLNPALAPVYTECLRLCGNWLAETCLESPAVILEK
YLERVVEVIEAESGVQDSRLLSQRTEAFLSLARFSDAQYQSIDKYMNSSEFENKQALLEK
AKEEVDLMKERKVTSNRYTIKVQREVELDEKALSNLQSDRQRFLCKAVENYIQCLEQGEE
HDTWVFRLASLWLENAVVKDVNDMMKKGLRQIPSYKFLPLMYQLAARMGTKMATGVAEDT
GFHDVLKELICRACLEHPHHTLFIIFALVNANKDENFCKTRVSKSTPKQPSPLDLERSDM
AQKIISAIRKKRGEMIRGIERLCDAYITLAYMDASRHKTEKKAIPIPADQPIMQIKDMDE
VVIPTMEIKVDPTGRYDNLVTIRSFMPQYHLAGGVNLPKIIDCVGSDGKSRRQLVKGQDD
LRQDAVMQQVFSMCSMLLQRNTDTRKRKLNIRRYKVVPFSQRSGVLEWCSGTVPIGEFLV
DPNKGAHKRFRPQDWTNLSCRKKMMEAQRLAFDEKLQAYSEVCKNFRPVFRYFCMERFLD
PAVWMEKRLAYTRSVATSSIVGYIVGLGDRHIQNILIDEQTAELVHIDLGVAFEQGKILP
TPETVPFRLSRDIVDGMGITGVEGVFRRCCEKTIEVMRSSQEALLTIVEVLLYDPLFDWT
MNPLKAFYLQHDEQQELNATLSSTMGGDDLVNHRKSSDSQSFNKVAERVLLRLQEKLKGV
EEGTVLSVGGQVNLLIQQAMDPENLSRLFPGWQAWV
NT seq
9291 nt
NT seq
+upstream
nt +downstream
nt
atgtgcagcacaatgaatttggctcttgatgatctccgggactgctgcagggggctggag
aatgacaaagccacagagaggaagaaagaagctgaacgcttcagacggctcattcgatgt
ccagaagttgtgcaggagttggatcgcacctcagggtctaaagctaaagcctccaagcaa
ctcacatgggatactgttttcagatttctgcagcgctacatccaaaaggagacagaaagc
atgcagtcaagtaaatccgtcacagcgacagcgctggccacaaggaagaagaagatggct
gaaatgtgcagtttgatcaaatacttcattcgttatgcaaacaaacgcgggcctcgtcta
aagtgcagtgagctgctgagacacgtgatggaggtgctgcaaagttcgtacagctgttca
gtttgtggagaagattatagcagtctcctagtaaaggacatcctgtccgtccgcaagtac
tggtgtgacatcgctccacagcagtggcagagtcttttggacttgtactgtagcttgttc
aactcttcatccaaacccatcaaccatgttttagtgagcagagtcatccacatagtggtg
cagggctgctgcatgcagactgatggatttaacaacactctgttcggcttcttctccaaa
gcacttcttaatgccagacaggagaagtacttgactgttttggagcacctcgtctcggct
ctcaacatattcttgagatctgcggccatgaactgtaggatgagggtgtgtcacctggga
gaggagcttctgccttctatactgcacatctgggtggataagaggcccagtgctgctctc
aaagagaagattgtggagttcttcaacctgcagatttgtgttcaccaccccaaaggagcc
aagacacaggacacaggtgctcatgctgaggactggacgaagtggcggagtcagctgtac
agcctttatgacgctttggtcagagaaatcagtcatataggcagcagagggaagtacgtc
acagggacaagacacatagccgtaaaagacaatctcattgagctcacagccgacatctgt
cgtcagctcttcactgatgacaatgagacccatatcttggaggtgacccaggcctccctc
agagccacgcagatgggcagtcccacccatggagcagccagcaagcgacgccgtatcgaa
ctgggctgggaggtcctccgggaccatctgcagtctcagcaaagtgactttgatatcata
ccatggctgcagatcactgcagtgcttgtctccaagtacccatccatggtgcctggccat
gaactggtcccactgctgtcagtgttgtaccagctcctggcagagcagcggagaggagag
agggggccatatgtgctgcgctctctgagggaggtggcccgctgccaggctcaccaccca
gagaggacccaggtccacagggccgaattgggcaggctatggggtcgggtgtgggccctg
gcactgcgaggggtcagctcaccccagactgaggccctcagcttcgacctgctggcctcc
attgtccatggtggacttattgatatggacagagagttctggaagcttttctctggatca
gcatgcaaactatcccagagtgccgctgtctgtctctcccaggccttgcttaagtgtccg
gtccccaagaggctcatcccgagctcaggttgggactatgtaggggtcacagaaggatct
gggtctccaaacctgaaggagaccctcatagcgtggctgctgatgagtgatcagagtgat
gagatggaggatagctccagacctcaccccatcatttgcagagactttccctacaatcta
atagcaggcatcctggtctccttgaccttgaaggacaccagagctggcctgaggtttctg
ctgggtgctgaaggagctaagagtgtcttttttcaagaacaagtgccatctgatgccaaa
gataacctggaggagatcgagaggctgtacttgcagttcagcttcaacacactgccctca
gaggtccgagggagcgaagagtcggtgaagccgtcatcagctgacggccaattcgctgcc
gtccctggtctcagaaacaagttggagcagtccctgctctgtgtggctgacaacctactc
aactgctactctcctgattctcagtccacccctccggagtgcctggtgcgctgcttcagt
ctgctgactggtgttctagcaggttacgtctccactggggttctgactgaagaggaggct
taccactcacaactcctcactaaggcaaaggccctggcccaggatttcagcgggtttgtt
tctactgtgaaagtaaagatgacagaggagggaaccatgcacacactgaagtctgtcatg
gagctctgcacacagtccgccagcagaaaaaacaaggataatgtttgtcccatctcctcc
agtctgttcattatgatactgccgggcagtctccttagtgaattggcagaggtcctcaaa
ctgttgctgagcagtgtttctaagagagtcagtttggtggatgaagacgatcatgcggat
gatgacagggatgtgaccaggactcaacaaggggacgatattgacctgtttgaggatggc
gaagaggctcacagcagcaccaatgagacacacaggcagaagggagacaacactgatgct
tcctgtgggccaggtgctaaaagcctgttagcagaggatcatctgggccagcaggacctg
gctctcctcgcgattttggagttcctatgtgagtgcggctctgtccagctcgtccatggt
ctgcttttcaaaccacaggaggtacagcgcaagctgctgctgctgttggagcagattgac
ttcaccaaagctctgcatctcaacatgtatcttgtcctgctgaagaagcttcctgctgag
gactcgctctctccagtggaatttgattcgctgctccgtcctctggcggacctgtgtaac
ctgtaccgtcaagaccaagagatctgtgttgctgtgctgctgggtttgttaccctcaatc
aggagcctggggaaaacccaccacatgcccgaggagatgaaacacgttcagggatccctg
ctacaagtggtgtctggattctgtttcctgagccagacagggaaatgcatatcaactatg
cgagccgccttggtgcaatgtctggttgctttgctggaggctgacccgtgttgtaagtgg
gcagtcctagatatcagagggtctgacaggatggaggaccgcctggtgtccgtcatcctt
ccgtcttaccttgcagatgcccaccaccaagtccggatgctggtagccatgtcagtggag
aggttgtttctggagatgagtccagagcagcctgataaaaagaagatgctgccacttaaa
caccagcaggatgcttttgaaaatgtttacctgaaggctcaggagggaatgaggctccca
aggagcagctcttcagaggacaagtgcgacgaggtgttcaaccgcaaggccaccctgctg
aagagtttgtctgtggtgttgtgttgtagcccagtgtgcgaaaaacagtctctgtttgcc
ctcttccagtcctacaaggagaacaatatcgaggaaaagctcattaaaaagatgttgtgc
agtgtatccagagtgctgggctacaggaatgtaaaaacatttgtcagttctcacttgtac
tacttagtggcagagtggctcgcacagagacaatccgacgatcgttacactctcggctct
ttcccctacaccctcctggaccatgacacagtcaaagatttctataactcttcctaccag
gtgttgatcccccacttggttttcctggacgactttgagcaggtgaggtctatcggcagg
tatctggagagggactggaaaaagctgttggccgactgcttccccaagatcatggtcaac
atcctgccgtacttcgccctgtctggccaggattcacaggtggcacagcagagagagaag
gctcatagagtgtatgacctgctcaaagatgccgactgtctgggcaaacagcaaattgac
agcctgatccacactaacctggcagacatagtagtggagctgctgatgactttgtatgaa
ggaggtgaagctgaaggagacagaggagacctgaaacgcttcataggggagctggaccct
gtaccaaacccaccatacttcagctcgtatgtcatcaaagccactctggactacctcagc
aaatgccacagtgccaaccacaagtcactggtagccattctgtcaaaaactccgatttct
atccaaaggatcctgctggcggtgtgtgaaagagcagcagagacaaccaacagctacgaa
cgtcatcgtatcctcctcatgtatcacctgttcgtcaacctgctgctcagagaggtgaag
gagggtctgggaggagcctgggccttcgtcctcagagacatcatctacacactcatacac
catatcaacggcagatcacttcagtgtgacgaggtttctacccgaagcctctctctgtgc
tgcgacctgttgacctctgtgtgtcaggcagccctgcagttctgtgacgacgctctagac
tgtcacctacaggtcattgttgggactctgacgtctcaggtgaccagccggcctgttatc
tcacagcaggtgctcagtctgttgcagtttctcgtgatagagaaccaacagaaattaaaa
ggagccatcagcaggttggaacctttccccgacctccctgaattcagagaactgcggtct
gtacagaacaggctcaaacataacactgggacctttacactgcgacaggaaatagtacac
ttcctgtcagtgtcgtcctgtgactccttacctctgaccagacttgagggactaaaggag
ctgacccgacagctgcatgacaacaagggacagatcagagagctgctcaaagagtgccac
gcggaccccagtgacagtgtgctggtgaagcttgttctgagtctgctgcagctctctaag
atagcagccaaccaccctggaggagccgacatactagaggcagcagggggctgtctagga
gaactgggtccagtagatttctccaccattgccctgctccatggcagagaccccttctat
gaaaaggcggtgtcactcttcacctccacagagttgcagtgtctttacatcattctgaac
tgcatgaacgacgcactgactcagccacgtattgaggtgaggcaggcagcagctcagtgt
gtgaagaacatcctcgccactcagtctggggttgacttctgggagcaacacaaagacaac
agagaccccatgctggcctaccttaatccctttagaaaggacaagagcaaggtggttgct
gtgagtgctgaggagagttcggaggccagggagaagttggagagtcaggaattttggatt
cctcaggcgggtggccacaaggcgtggctgagggctctgtgcaccgccctgctggatagt
ggaggagtcaagagcgaggctctgctgctgtcgcgaccactgtgtctggtgaaggtggac
tgctgtcagaggttgctgcccctcatcatccactccatccttctagatgactctgatggt
tcgtggagggagttgctctcatcccacatccaggacttcttcagtttctgttccagaagt
gctcaggcctccagtcggtctgctacacctctcaattctgactcagattctgacacagcc
agccagggcttgtatgacaagacctctctgcgtaccatgctggcagttattgactatttg
agacaccaacagagaccactggactctaatagtaagtcctgcggcacggtgtgcaactca
aacttctggctggagctgaactatctggaggtagccaaagctgctcagtcctgtgcagct
cacttcaccgctctgctgtatactgagatctacgtcgataagatcaaagctaacatggag
gaaagtcgcagaaccaagtccagagcgacatataagctcagttttgaggagcacagccag
aagttcaccatctccagcctgacagagaaaagtatggaagacaccaatatcagtctgcag
gagcttcttattgaggtgtaccggagcattggagaacctgatagtctgtatggctgtggt
ggggagacgatgaccagttcactgaccaggattcgaacctacgagcacgaggctatgtgg
gggaaggccctgacctcatatgaccttcactccactctgcctgaggccactcgccaagtg
ggaattgtggtgggcttgcagaactttggtctgagcagcatcctcgccacctacatgagg
ggtctggagagtgaaggggtggagtggggggctgagctaagagagctgcgcttccagtcc
gcttggaggaacacccagtgggactgtgagctgtcagagaggagtgacaaatccacacct
ggcttccatgaatcggtgttttgttccctgcaagcgctgaggaacaaagagttctccata
ttcgatgaaacgctgaaacaggccaggggtgcagaggtggaggagctatgcagaggcagt
ctggaggctgtgtcttctctgtacccggctctcaggaacctgcagagcatcagggagctg
gagagtgtgaaacagctcttctctagacccttctcagatgtgactctgagggatgtttgc
agccagtggcggcagcactcccagctgctcgctgacagtgactttgctttggtggagcca
attctgaccgtccgctctgtggcccagaacaccctgatgtccagggtgggagaccccgac
agcatccagtacctcagctctgtgctcactgatcacctcatggagctctgccagctggct
cgcaaggctgggaatacacagcgagcagagcaggcagtcttccggatgaagcagcatggt
ggcggagggtcctgggcgtcgtcacctgtattgtcatggcaactggaggaggcccaggtg
ttctgggcaaagggagagcagggtctggcactaggcctgctgagacagatgatacacaac
ttagagaaaagggtggatttaaatcctgctctggctccagtctacactgagtgtctgagg
ctgtgtggcaactggctagctgagacctgcctcgaaagtcctgcagtcatattggagaag
tacttggaaagggtggtggaggtgatcgaggcagagtcaggagtgcaggactccaggctg
ctgagtcagcggaccgaggccttcctgtctctggcccgcttctctgacgctcagtaccag
agcatcgacaaatacatgaactcctctgagtttgagaacaagcaggcgctgctggagaag
gccaaagaggaagtggacctgatgaaggagcgtaaagtgacctccaacaggtacacaata
aaggtgcaaagagaggtggagctggatgagaaggccctgtccaacctgcagtcagacaga
cagcgctttctgtgtaaggcggtggagaattacatccagtgtctggagcagggcgaggag
cacgacacctgggtgttccgcctggcttcactgtggttggagaatgctgttgttaaggat
gtgaatgacatgatgaagaaagggctgaggcagatcccctcctacaagtttctgcctctc
atgtatcagctggctgcacgcatggggaccaagatggcaactggcgttgctgaggatacg
ggcttccatgatgtcctcaaggagctgatctgccgagcgtgtctggagcaccctcaccac
acgctcttcatcatcttcgccctggtgaacgccaacaaagatgagaacttctgcaagacc
cgggtgtcgaagagcactccaaagcagccatcaccacttgacctggagcgttcagacatg
gcccaaaagatcatcagtgcaatcaggaaaaaaagaggcgagatgatccgagggatcgag
cgtctgtgtgacgcctacatcactctggcttatatggacgccagcaggcacaagactgag
aagaaggctattcccatccctgctgatcagcccatcatgcaaatcaaagacatggatgag
gttgtcatccccacaatggagatcaaggtggacccaactggtcgctatgacaacctggtg
actatcaggtccttcatgcctcagtatcaccttgccggaggagtcaacctgcccaagatc
atcgactgtgttggctctgacggcaagagcaggagacaactggtcaagggtcaggatgac
ctgcggcaggatgcggtgatgcagcaggtcttcagcatgtgctccatgctgctgcagcgc
aacacagacacgcgcaagaggaagctcaacatcagacgatacaaggtggtgccgttctcg
cagcgtagcggcgtgttagagtggtgctcaggcactgttcccattggggaatttctggtg
gatcccaataaaggagcccacaaacgcttccgaccccaggactggaccaacttatcctgc
cgcaagaagatgatggaggctcagaggcttgcatttgacgagaagctccaggcttactct
gaggtgtgcaagaacttcaggccggtcttcaggtatttctgcatggaacgattcctggat
ccagcggtgtggatggagaaacggctggcttacacccgcagcgtggccacctcctctatt
gttggttacatcgtaggtctcggcgacagacacatccagaacatcctgattgatgagcag
actgctgagctggtccacatcgatttaggtgtggcctttgagcaggggaagatcctcccc
acaccagagactgtgcccttcaggctctccagagacatcgtggatggaatgggcatcact
ggagtggagggagttttcaggagatgttgtgagaagaccattgaggtgatgaggagctct
caagaagctctgctgaccattgtagaggtgctgctgtacgaccctctgtttgactggacc
atgaaccctctgaaggccttctacctgcagcacgatgagcagcaggaactcaatgcaaca
ctgagctccaccatgggaggagacgacctagtcaaccaccgtaaatccagtgacagtcaa
agtttcaacaaggtggcagagcgggtgctgctgcggctgcaggagaaactgaagggggtg
gaggagggcacggtgctcagcgtgggggggcaggtcaacctgctcatccaacaagccatg
gaccccgaaaacctcagcagactctttccaggctggcaggcctgggtgtag
DBGET
integrated database retrieval system
