Tetraodon nigroviridis (spotted green pufferfish): GSTEN00015888G001
Help
Entry
GSTEN00015888G001 CDS
T04346
Name
(GenBank) unnamed protein product
KO
K08809
striated muscle-specific serine/threonine protein kinase [EC:
2.7.11.1
]
Organism
tng
Tetraodon nigroviridis (spotted green pufferfish)
Brite
KEGG Orthology (KO) [BR:
tng00001
]
09180 Brite Hierarchies
09181 Protein families: metabolism
01001 Protein kinases [BR:
tng01001
]
GSTEN00015888G001
Enzymes [BR:
tng01000
]
2. Transferases
2.7 Transferring phosphorus-containing groups
2.7.11 Protein-serine/threonine kinases
2.7.11.1 non-specific serine/threonine protein kinase
GSTEN00015888G001
Protein kinases [BR:
tng01001
]
Serine/threonine kinases: CAMK group
Trio family [OT]
GSTEN00015888G001
BRITE hierarchy
SSDB
Ortholog
Paralog
GFIT
Motif
Pfam:
I-set
Ig_3
Pkinase
Ig_2
ig
PK_Tyr_Ser-Thr
V-set
C2-set_2
fn3
Ig_DSCAM-DSCAML_C
Pkinase_fungal
C2-set_3
ABC1
Receptor_2B4
Ig_4
Izumo-Ig
Kinase-like
APH
Motif
Other DBs
NCBI-ProteinID:
CAF98242
UniProt:
Q4SM90
LinkDB
All DBs
Position
3
AA seq
3248 aa
AA seq
DB search
MAESPPPGPSPNVPPKRPKVTADPAVPHPGVGSQESQPPAFIRKLRKAAVGTGCDIRLRV
SVAGFPKPRLSWYHNDDLLPPSEAQDPGGLWIRDCRTSDAGLYTCVATNQMGEASCSAVL
AVMDLGEVSHSGALSPRRTCHPGSLEDLEFRKLGIPHGHLADSPMVCYSETTEDETTEPQ
VETKEGSGRHQDKAGWRRASGEGGGMMDYNPDASDRRATLPGAYDPVVERELRALGSRPP
GLHPDPSDPARQALGGAPAEGGPPVRHLGVEPLIRASHANLARPVQGTYSSCSSALVTSH
LHFPLNPHPSRTQTAEESQETSPHDPLFSPRPPANSHPNSIPNRDLDKDSPSDHRSETDQ
LSNANLDHQEPKISGKIRAFENRIASSEKPGGSAGGRVASPDSSDGVRKTGGPSREEARI
LQGAAQKRATFKQRASSLEDKTNYSQRVQSYQSKFTEELQRIKKLVGKPSLKKAFSSEQL
SQKDRLAAEKLEPIPPQVVQKLEARERAMREREAREREGKSQALLQGYKTKRSLHQQRYI
RCRTNKSGHQRRTFSSSQTRIIQSAWKTSFPFNGGGSWSPEATQDPPDPITRPIRKPPAQ
KKEGRDMSPCESEHPNNPGGTYLSADGEMAEGPRFQTLLEDKVAPSDTAVVLKCVVTGRP
PPTVTWRKDNVAIRSDAFHVVASEGDSHSLLIKQMRPSSVGSYCVTAVNPAGTASCSATV
HVQPELACSPSLSAEEAMELGDSPASWKKVHFKEPPSFQVVPFDQAVTEVQDVVISAKVS
GQPTPMVHWSKDRVPVKTAGRFAVQQTDNGTLELRISSAQRSDAGLYACRIINESGSRQA
ECRVDVRGAGDTTLTITREVADVAVKAGESVMLECHMAGPSDVDVDWLSNGKLIQPALLN
CKMHFDGKRCRLLLNSVHEDDSGTYTCKLSTAKEELTSSANLKVTPSREPLFTRKLDVLE
VTEGRSARFDCKVSGSPAPRVTWMHFEARVEEGENAHILQAGGRHSLVIAHVSNDSEGFY
TAIAQNIHGQSECTAELYVQEHRAAISSHMAKMEKMPSIPEEPEVLESEVERRTMPDFVK
PLADMEVVEGKEVALRCKVAGLPYPTISWYHNSRRIESSEERKMTQQRDVHSLVIRSACH
AHGGVYKAVISNKVGKAACYAHLYVTDIVPEAPDGAPVIEAITGKTITISWKKPKKLDPS
LDPGSLLYVVQQQPLGSIQWSVVASNLKETSYTITSLSKGVRYAFRVLASTGKTLSKPSL
STDLIQLLDRGPYLRKAPIILEKPDCVYVVENQPASITITLNHVHAVVTWKRRGVVLASK
PGMYEMSMPDDDQHMLKLQRVRSTDVGQLVVMASNQFGSDLCALQLALAVRPKFETIMED
VDVYVGETTRLAVVVEGKPDPDILWYKDDVLLKESSHFTFVYDDPEYSLVILNACAEHSG
VYTCTAKNLAGTNSCKAELTVRTERKEEAEPMEDEGTILRKMRRLVDYYDVHKEIGRGAF
SYVKRVTQKKAKAEFAAKFICARGKRKASALREMELLSELDHERILYFHDAFEKKNVVVL
ITDLCHEELLERMAKKTAVKELEVSLFASPALRHPYVTLSFTLVFPQIRWSIQQVLEGLW
YLHQKNIAHLDIKPENILMASPTSDQIRICDFGNAAKLDPSEEYYCKLGTPEFVAPEIVN
QTPVSTATDIWPVGVIAYLCLTGISPFAGENDRATALNIRNYNVAFEETMFSDLCREARG
FVIKLLVVDRLRPSAAECLRHPWFKVSVRGQGINTALLKQVLSRRRWQRSLLSYKSKMVM
RTIPELLNDSSNHVSIAVARHLKEGSPPPSSSSDSDADVDELPFIPMPLSMFFSSSRVSL
NEIPGDEDVTASQMSRNTDDTGAVAEKDEEALKEENMEKTKRAPLKKGSSVETEDSETRA
RRATMRRGSSADSALLIHVESEEENSSSSEATNKSLKKAVSMELPHRSPSPGLAKISQEE
YALKLELMRQRLLRGGNDKKMSGLRGPLFETLGIEDDRRTGSLDRNLRRSRTGPPSLARA
ASSESPAEDRPLTKVFCKSASFNQGDSEPMPLHRRYGAPLEIPAVGNGNTEGKKLQEATS
MSALTEQATPESASPTKKTSFSLPALAEEDDEDEEKKEMQELYINEKNFQDEKETRNSSS
VSLAPLALLSNKAESRPNRVDTGKPNGMTSTNGATRPEHAAVFSKVATTLRPSAPSSSTA
PRQPVLRTDIKNIDSEEIFEARFKKRESSLTRGLRKLTRNKSEEKSPTLSRKGAGGAEEV
YRPGPKGAPLEMVSRGLQEKSKSVQDLRGEDKEPGLGLIGRFSLRAKRSSSIDKKGEKPK
EEKPPDIQESAVNKRVSWAVGRSKSLDTNEPSSTKRLQGEREERNAESSVSAMRRKFESK
VAGMSAKVRTQSEERKDNDAAGCQKDPKRLTDSPITAIRHKFESKVAGLSSKIRSQSEER
KDDPDERRTPLFARHRHSHSEGRGLKGMGIPENQLAKQTGTNASKDSLDSTSSLQSEKVP
ESDRRSRWDRWGLTRSRKDKSPSQPDLSSSSRKDEQQFIRAASDFAPVFHIKLKDHVLSE
GEPATLSCLPAGSPPPRITWRKDRKLLEVDDRMSLISHPDGRQLLQIAKSNPKDDGVYEC
VASNPIATITTSCTLSVASVPKRPGTPEVAQTYNNTALVLWKPADTKSPCTYTLDRKTDG
DSDWMTVATGVTDCYYNVTELPPGGTFRFRVTCSNKAGQGPASNSSGPVSLDPAGGRALP
AATVGKTLPFPLEPVPPPKPVQNAPGTVNSCVSPPKSGAPPPSRTEVASSAPTQQNRDTQ
HFAASPPIKIPPPLITPKPQSPVNVVCPMTQSPPPAVTPPLTPTKPTVPTYAPVPVTSAC
VAPTPVIFSPQVLQSSSLSPFADGVTTPTKGTPSGRATPSTGLRQGVPQETLHLPGREGQ
GSLRRCPRVPGKRHGQNLPGQDRPLQPGEQTGGPEGVRDPESLHSEKIMALHEAYVTPRY
LVLVAEFCSGKELLHSLIDRFRYSEDDVVGYLVQILQGVEYLHSRRVLHLDLKPDNIMVT
NLNTIKIVDFGSAQSFNPLSLKQKDARTGSLEYMAPEMIKGEVVGPPADIWSIGVVTYIM
LSGRLPFEDQDPRQVESKILVAKFDPSKLYPNVSQSASAFCKKMLSSYPWARPATRDCFA
QAWLQDSYLMKLKRQTLTFTSNRLKEFLVEQQSRRSEGATKHKVLLRTYQSSPQSPGAGS
TLHSPAPQ
NT seq
9747 nt
NT seq
+upstream
nt +downstream
nt
atggcagaatctcccccacccggtcccagtccgaacgtcccacccaagagacccaaagtc
acggcggatccagcggttcctcatcctggggttggatcacaggagtcccagccgccggcg
ttcatccgcaagctccgcaaggcggcagtggggacgggttgtgacatcagactgagggtc
tctgtggctggatttcccaaacctcgactcagctggtaccacaacgatgacctccttccc
ccctcagaggcccaggacccaggggggctctggatccgtgactgccgcaccagcgacgcc
ggcttgtacacctgcgtggcgaccaatcagatgggcgaagctagctgcagcgccgtcttg
gctgtgatggaccttggtgaagtgagtcattctggggctttgtccccgcggcggacctgc
catccaggaagtttagaggatttagaattcagaaaactgggcatcccacacggacatttg
gcagattcacccatggtgtgttactctgaaaccacggaggatgagaccacagagcctcag
gtggaaaccaaggaaggttccgggagacaccaggacaaagctggatggagaagagcttca
ggtgagggaggagggatgatggactacaacccggatgcctcggaccgtagggcaactctt
cccggtgcctacgaccccgtggtggagcgagagctccgcgctctgggctcccgccctccc
gggctccacccggacccctcagacccagccaggcaagccctcggtggcgctcctgccgag
gggggccctcctgtccgtcacctgggtgtcgaaccgctgatccgagcttctcacgctaac
ctggcccgacccgttcaaggcacctacagcagctgctctagtgctctagtaacgtcacac
ctccatttccccctcaacccccatccctctaggactcaaactgcagaggaatcacaggag
acctctccccatgaccccttgttctctccccggccacctgccaattctcaccccaacagt
atcccaaaccgagacctggacaaggacagcccttctgatcacaggtctgaaacagatcag
ctttcaaatgccaacctggatcaccaggagcctaagatctctggcaaaatccgagcattt
gaaaaccgaatagcaagttcggagaagcctggcggatctgctggaggtcgagtagcttct
cctgattcaagcgatggggtgagaaaaaccggtggccccagcagagaggaggccagaatc
ctgcagggtgcagcccagaaaagagccaccttcaaacaacgagcttcctcattggaggac
aaaaccaattattcccagagggtccagagctaccagagcaagtttacagaagagttacag
agaatcaagaaacttgtgggaaagccgagcttgaagaaggctttttcctccgagcaactt
tcacagaaggacaggctggctgcggagaagctggagcccattccgccccaggtggttcag
aagttggaggcaagagagcgagctatgcgggagagggaagccagggaaagggaagggaag
tcgcaggcgctcctgcagggatacaaaaccaaacgttcactccaccaacaaagatacatt
cgctgccgcacgaacaaaagcggccatcagagacgcacattcagcagcagccaaacccgg
atcatccagtccgcatggaaaacgagcttcccctttaacggtgggggaagctggagccca
gaggctacccaagaccctccagaccccatcacccgtcccatccgtaagccccctgctcaa
aagaaggaaggcagagacatgtccccctgtgagagtgagcatcccaacaatcctggtggt
acgtacctgtctgctgacggggagatggcagagggccccaggttccagacgctattggaa
gacaaggtggcgcccagcgacacggctgttgtactcaaatgtgttgttaccgggaggccg
ccacctacagtaacgtggagaaaggataatgtggccatacggagcgatgctttccacgtg
gttgcgtccgagggggacagtcacagcctgctgataaagcagatgaggccgagcagcgtt
gggtcgtactgcgtcacagccgtcaacccggcggggacggcgtcctgcagcgccaccgtc
cacgtccagccagagctggcatgcagtccgtctctgtcagccgaggaggccatggagctc
ggggactcgcctgcttcctggaaaaaagtccactttaaagagccgccctcatttcaggtg
gtgccgttcgaccaagcagttactgaagttcaagatgtggtcatctcggcgaaggtcagc
gggcagcccacacccatggtgcactggtcgaaggacagagtcccggtgaagacggcgggg
cggttcgctgtgcagcagacggacaacggcaccctggaattgagaatcagctcggctcag
aggtcagatgcgggactctacgcctgcagaataatcaatgaatctgggagcaggcaggcg
gagtgcagagtggacgtcagaggtgcaggagacacgactctgacaatcaccagggaggtg
gccgacgtggcggtgaaggctggcgagtcggtcatgctggagtgtcacatggcaggaccc
tcagacgtggatgtggactggctgtcgaatgggaagttgatccagccggcgctgctcaac
tgcaaaatgcactttgatggaaagaggtgccgcctgctgctgaattcagtgcacgaggat
gacagtggaacgtacacatgcaagctcagcacagccaaagaggagttgacctcgagtgca
aacctgaaggttacgccctccagggaacccctgttcacccgtaaactggacgttctggag
gtcaccgaaggccgcagcgcccggtttgactgtaaggtgagcggctcacctgctccgcga
gttacttggatgcactttgaagccagagtggaggagggggagaacgcccacatccttcaa
gccggcggtcgtcactccctcgtcatcgcccacgttagcaacgactctgagggcttctac
accgccatcgctcagaatatccacggacagtctgaatgcaccgccgagctttacgtccaa
gaacatcgagccgctatctcctctcacatggccaaaatggagaagatgccatccattcca
gaagagccagaggttctggagagcgaagtggagcgacgaaccatgccagactttgtcaaa
ccgctggctgacatggaggtggtcgaaggcaaagaagtggcgctgaggtgcaaggtggct
ggcctcccgtacccgaccatcagctggtaccacaacagcagacgcatcgagagcagcgaa
gagcgcaaaatgacccaacaaagggatgtccacagtctggtcatcaggagcgcttgtcac
gctcacggaggcgtctacaaggctgtcatctccaacaaagtgggcaaagcagcttgctat
gcccacctctacgtcacagacatcgtccctgaagctcctgatggtgctccagtgatcgag
gccatcacagggaaaacaataaccatcagctggaagaagccaaagaagcttgacccttct
ctggatcctggctccttgctctacgtggttcagcagcagccactgggctccatccagtgg
tctgtggtggcctctaacctgaaggagaccagctacaccatcacctccttgtccaaggga
gtacgctatgctttcagagtgctggcgtccactgggaagaccctgagcaaaccgtccctg
tccacagacctgatccagctactagacagagggccctatctaagaaaagcaccaatcatt
ctggagaaacccgattgtgtctacgtggtggagaaccaaccggcgagcatcactatcacg
ctgaaccacgtgcatgctgttgtcacctggaaacgacggggggtcgtgttggcgagcaag
ccggggatgtatgagatgagcatgccagacgatgaccaacacatgctgaaactccagcga
gtacgcagcaccgacgtgggccagttggtggtcatggcaagcaaccaatttggcagcgac
ctctgcgcccttcagctggccctggcagtgcgaccaaagtttgaaacaatcatggaggat
gtggatgtgtatgtaggggagacgacccgcttggccgttgtggtggaagggaagcctgac
ccagacattctgtggtataaggacgatgttctcctcaaagagagcagccacttcaccttc
gtctacgatgacccagaatattctctggtcatcctcaatgcctgcgctgaacactccggc
gtgtacacctgcactgcaaagaacctggccggcaccaactcctgcaaggcagagctgacg
gtccgcacggagagaaaggaagaggccgagccgatggaggatgagggaaccattctgagg
aagatgaggcgcctggtggattactacgacgttcacaaggagatcgggcggggtgccttc
tcctatgtgaagagggtgactcagaaaaaggcaaaagcagaatttgccgccaagttcatc
tgcgcgcggggcaagaggaaagcctcggctctccgagagatggagctgctgtctgagctg
gaccatgagaggatcctttatttccacgacgctttcgagaagaagaacgtggtggtacta
atcacagatctatgtcacgaggagctgctggagcgaatggccaagaaaacagcagtcaaa
gagctggaggtgagcctgtttgcgtcgcccgcactgcgccacccttacgtaacgctgagc
ttcacacttgtttttcctcagatccgctggagcatccagcaggttttggaaggtctgtgg
taccttcaccagaaaaacatcgcccaccttgatataaagcccgaaaatattttgatggca
agtcccaccagtgaccagatccgcatctgcgacttcgggaacgcggccaaactggaccca
tcggaagagtactactgcaaactcggcacgccggaatttgttgccccggagattgtgaat
caaactcctgtctccacggcaacagacatttggcctgtcggcgtcatcgcttacctctgc
ctgaccggcatttccccttttgctggggagaacgacagagccacagccttgaacattagg
aactacaacgtggccttcgaggagaccatgttttctgacctctgcagagaagcgaggggc
ttcgtcatcaagctgctggtggtggacaggctgaggcccagcgccgctgaatgccttcgc
catccttggtttaaggtgagtgttcgaggtcagggcatcaacacggcgctgctaaagcag
gtcttgtctcgtcggcgatggcagcgatctcttctcagctataaatccaagatggtgatg
cgaacgattccagagcttctgaatgactcatccaatcatgtctctatcgccgtggctcga
catttaaaagagggctctccaccaccctcctcttcctctgattcagatgcagatgtcgac
gagcttcctttcatccccatgccactctctatgtttttttccagttcaagggtctccctt
aatgagatccctggagatgaagatgtcaccgcatcacaaatgtcacgaaacacagacgac
accggggctgtggcggaaaaagatgaagaagctctaaaggaggaaaatatggaaaaaact
aaacgagcacccctcaagaaaggatcgagtgtggagacagaggattccgagacaagagcc
agacgggccactatgaggagaggcagctctgcggactcagccttgctgatccacgttgaa
tctgaagaggaaaattccagcagttcagaagcgactaacaagagcctgaaaaaggccgtt
tccatggagctccctcatcgcagcccgagccctggcctggcaaagataagccaggaggag
tatgctctaaaactggaactcatgaggcagcgcttgctcaggggaggaaatgataagaag
atgagcggtctgcgtgggccgttgtttgagacgcttggcatagaagatgacaggaggacg
ggatcccttgatcggaatctgaggagatccagaactgggccgccatcgctggccagagca
gcatcctctgaaagtccggcggaggacaggccactgaccaaagttttttgtaaaagtgcc
tcctttaatcagggcgattctgagccaatgcccctgcaccgcaggtatggagccccctta
gaaatcccagcagtcggaaatgggaacactgaaggtaagaagctccaagaggcaacatcg
atgtcggcacttacagagcaagccacaccggaatctgcctcgcctacaaaaaagacctct
ttcagcctcccagcgctcgcagaggaagacgatgaagatgaggagaagaaagaaatgcag
gagttgtatataaatgaaaagaactttcaggatgagaaagaaactagaaacagtagtagt
gtgtcattggcacctctggctctgctgtctaataaggctgaatccagaccaaaccgtgtg
gacactgggaagcccaacggcatgacttccaccaacggcgccactcgtcctgaacatgca
gcggtgttttccaaggtcgccaccacgctccggccttccgcaccgtccagttccactgcc
cctcgccagcctgtgctgaggacagacatcaaaaacatcgactcagaggagatctttgaa
gcccgcttcaagaagcgtgagtcgtctttaacccgaggcctccgaaaactgaccaggaat
aaatcagaggagaaatcaccaacactgagccgaaagggagcagggggggcggaggaggtt
tacaggcctgggccgaagggggcacctctggaaatggtatccagaggattgcaggagaaa
tctaaatctgttcaagacttgagaggagaggataaggagccaggccttggtctcatcgga
aggttttccctgcgagccaagaggtcgtcgtcaatagacaagaagggagagaagccgaag
gaagagaagcctccggacattcaggaatccgcagtcaacaagagagtttcctgggctgtc
ggtcgcagcaagtctctggatacgaacgagccaagcagcacgaaaagactacagggggaa
agagaagaaagaaatgccgagtcgtccgtttctgccatgaggcgtaagtttgagtcgaag
gtggcaggaatgtctgccaaagtgaggacccagtcagaggagaggaaggataacgatgct
gcgggatgccagaaagatcccaagagacttaccgattcgcccatcacggccatacgacac
aagtttgagagcaaagtagcaggattatcttccaagatacgcagtcagtctgaagagagg
aaggacgatccggacgaaagaaggacgcctctgttcgctcgccatcgccactcccactca
gagggacggggactgaagggaatgggaatccctgagaaccaactggctaagcagacaggc
accaatgcatccaaggactcgcttgactcgacgtccagcctccagtctgaaaaagtcccg
gagagcgacaggaggtcccggtgggacaggtggggtttgaccagaagcaggaaagacaag
agcccgtcccagcctgacctgtcatcatccagccgcaaagacgagcagcagtttatccga
gcggcttccgattttgcccccgtgttccacatcaagctgaaggaccacgtcttatccgag
ggagaacctgcgactctgagctgcctcccagctggaagtccacctcccagaattacatgg
aggaaagatcggaagctgctggaggtggatgacagaatgagcctcatatcccacccagat
ggccggcagctccttcagatcgccaagtccaacccgaaggatgacggcgtttacgaatgc
gtggctagcaaccccatcgctactatcaccacctcctgcacgctgtcagtggcttctgtt
cccaaacgtccagggaccccagaagttgcccagacatacaacaacacagccctggtgctc
tggaagccagcagacaccaagtctccctgcacctacaccctggacagaaagacagatggt
gactctgactggatgactgtggcaactggagttactgactgttactacaacgtcacagag
ctaccgccaggtggcacctttaggttccgcgtcacctgcagcaacaaggcaggacaggga
cccgcgagcaacagttccggacctgtgagcctggacccggcaggtggcagagctttgcct
gccgcaacagtgggaaagacacttccatttccactggagccagtgcctccacccaaacca
gtccagaatgctcctgggacggtaaattcctgtgtctcccctccaaaaagtggagcccca
cctccgtctaggacggaagttgcttcgtctgcacctacgcaacaaaaccgggacactcag
catttcgccgcctcgcccccaatcaaaatccctccacctctcatcacgcccaaaccacag
agtccagtcaacgtggtgtgcccgatgacccaaagcccacctcctgctgttaccccgcca
ttaacaccaaccaagcctacggtgcccacctacgcccccgtccccgtcacctccgcctgc
gtggccccgacccccgtcatcttctccccacaggtgctccagagctccagcctgagcccc
tttgcggatggggtgaccacgcccaccaaggggaccccatccgggcgggccacgccttcc
actggtctgcgccagggggttccacaggaaaccctacaccttcctggacgagaaggccag
gggtcgcttcggcgttgtccgagagtgccgggaaaacgccacgggcaaaatctacctggc
caagatcgtcccctacagccaggagagcaaacaggaggtcctgaaggagtacgagatcct
gaaagccttcacagcgagaagatcatggcgctgcatgaagcgtacgtgacgccgcgctac
ctggtgctggtagccgagttctgcagcggcaaagagctgcttcacagcctcattgacagg
ttccgctactctgaggatgacgtggtgggctaccttgtgcagatcttgcagggagtggag
tacctccacagccggcgtgtcctccacctggacctgaagcccgacaacatcatggtgacc
aacctcaacaccatcaagatcgtggactttggaagcgctcagagcttcaacccgctcagc
ctcaagcagaaggacgccaggacaggatccctggagtacatggctcctgagatgatcaaa
ggtgaagtggttggtcctcctgcggatatttggtccattggtgttgttacctacatcatg
ctcagtggccggctgccctttgaagatcaagatcctcgacaggtcgagtccaaaatcctg
gtggcaaagtttgacccaagtaaactctacccaaacgtgtctcagagcgcctccgccttc
tgcaagaagatgctgagcagctacccctgggcccgcccggcgacccgagactgcttcgcc
caggcctggctgcaggactcctacctgatgaagctcaagcgtcagactctcaccttcacc
tccaaccggctcaaggagttcctggtggagcaacagtcccgccgctccgagggcgccacc
aagcacaaggtgctcctgcgcacctaccagagctcgccccagtccccgggggccggctcc
acgctgcacagccccgccccccagtga
DBGET
integrated database retrieval system