Tsukamurella paurometabola: Tpau_1315
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Entry
Tpau_1315 CDS
T01228
Name
(GenBank) MaoC domain protein dehydratase
KO
K11533
fatty acid synthase, bacteria type [EC:2.3.1.-]
Organism
tpr
Tsukamurella paurometabola
Pathway
tpr00061
Fatty acid biosynthesis
tpr00074
Mycolic acid biosynthesis
tpr01100
Metabolic pathways
tpr01212
Fatty acid metabolism
Module
tpr_M00083
Fatty acid biosynthesis, elongation
Brite
KEGG Orthology (KO) [BR:
tpr00001
]
09100 Metabolism
09103 Lipid metabolism
00061 Fatty acid biosynthesis
Tpau_1315
00074 Mycolic acid biosynthesis
Tpau_1315
09180 Brite Hierarchies
09181 Protein families: metabolism
01004 Lipid biosynthesis proteins [BR:
tpr01004
]
Tpau_1315
Lipid biosynthesis proteins [BR:
tpr01004
]
Fatty acid synthase
Bacteria type
Tpau_1315
BRITE hierarchy
SSDB
Ortholog
Paralog
Gene cluster
GFIT
Motif
Pfam:
Fas1-AflB-like_hel
Acyl_transf_1
FAS_AT_central
MaoC_dehydratas
ketoacyl-synt
Ketoacyl-synt_C
DNA_pol_B_N
SAT
adh_short
Fas_alpha_ACP
Patatin
FAS_I_H
adh_short_C2
Motif
Other DBs
NCBI-ProteinID:
ADG77944
UniProt:
D5UWS2
LinkDB
All DBs
Position
1361005..1370313
Genome browser
AA seq
3102 aa
AA seq
DB search
MTVDDFVTAHARTAKNPRSNDALVDRLASGEQYALIFGGQGGPWLAGLVSLLEQTGLADD
PEYRELFDGFVSRADSALEPVRAELLRAQALTFDPMRWRDASIAASEAETTDEEGDEPAQ
AATQSRRPLAPQVDGAVFSVPGVLFSQLVALYGLEAQGLDPWDVPPVAAIGHSQGQLAVA
AFASRGAEGDLLAIARLIGAAGRVVARRKGLVALGGATPMMSVSGVRQDKLEQLLADFTA
TVAFDACGPVLAVRNSRTAFVLTGTPKLLNGFREYCESVAETEEAQRKDKLTGGAAFAPK
FEDVPVDIGFHHPALADGVALVTEWAERCGLDAEKAGRLAAAVLVEPVDWVADVDAAIAK
GARWLIDLGPEDLATRLAAAPARGRGVGLVPATTLRGARNLFSPGGIPELPLAWSEFGPR
LVALPDGRTVLSTRFTELTGRSPILLAGMTPTTVDPGIVAAAANAGHWAELAGGGQVTEP
IFADNIAKLTALLEPGRAAQFNSLFLDPYLWKLQLGGKRIVQKARAAGAPLDGVIITAGI
PELEEATALVEELTDAGMRYVAFKPGTVEQIRQVARIAAEVPRYPVIVHVEGGKAGGHHS
WEDLDELLLSTYAELRDRGNVVLAVGGGIGTPELAAEYLSGEWSRKYGAPAMPVDGILVG
TAAMATLEAATSDDVKRMLAEIPGTPDWIGAGHAGGGMASGRSQLGADIHEIDNTASACG
RLLDEVAGDGDAVAERRDEIVAAMAGTAKPFFGDVDAMTYAQWLERYLELSVGTRAQVAA
ADPERFGDYGWLDITHQDRFLSMLHRAEARLAAEDRGPIETAFADVTATDDAYAAFDALL
QRYPDAASTRLHPADVPFFVSLCRTPGKPVNFVPVIDKDVRRWWRSDSLWQAHSPRYTAD
EVCIIPGPEAVAGITRVDEPVGELLNRFEAELVRRRTNAGERAEAVAGRRVVPGVESVPL
ARALAATNVLWAGRLVHSPIARLGAGSTWAIHGEGDESVALQPDTGARVRVTGPNSVRLE
IPVSGTELVIPIEVPDSCADGGVPVIELGEASEAMSALVALAAAGDLPAVHDGAATIDFD
FRAEDAADHAAITADSLPVGLSALYGDEQLAVPDAFVGLCWPASFAVVGAAKTAEGRPVV
EGMLDLVHLDHAIELAANPAPGSYRATATLEDVTATDIGTVVTVGIDVRAAGTDAPAVTM
TERFAIRGRTGEAELADPQPAGGARGETATETPRKRVRQVTVQAPQDMSGFAAVSGDHNP
IHTSQVAASLAGLPGPIVHGMWLSAAAQQVTSASNGDVMHRKLRGWTTRFLGMVLPGDEV
QFTVERVGIDRGGELLEISARVGENLVMAASAVAFPAVTAYVFPGQGIQGKGMGLAQRSR
SKAAREVWDAADEHTRENLGFSILAVVRDNPLSIRANGTVYTHPDGVLYLTEFTQVAMAT
VACAQLAELTEIGARVDDPIIAGHSVGEYNALASAGVLPLGTILETVFHRGRIMQQLVPR
DAEGRSPYAMAAIRPELFEVDDADLANWVEGSLEGADEYAEIVNLNLRGAQYVVVGNQAG
LDALEAAIAERTGSPKAFQLVPGIDIPFHSSVLRAGVPEFAARLEELVPAEFDYTKLVGK
YLPNLVARPFELTADFALSILDVVPSEPVRELVADWERAAANPMATARTLLIELLAWQFA
SPVRWIETQDQLLIDTDHGGLGVERFIEVGLASAPTLANLASRTLALASYQRAAATVLNA
ERDQAKVFATDEDAPEDEADDAGTEEASAAPVSVAAVAPSAAPAAPAGGPRPDDLSFDAS
DATRAMIAIWTKMRPDQLEKTDTIETLCDGVSSRRNQLLLDLGSELNLGAIDGAAEADLP
TLAGTVTKLARTYKPLGPVLSETVGNQLRKLLGSTGKRPNYITEYVTGTWELGPGWAQHT
VLALATGTREGASVRGGDLATLLDGPVSNAAALDALIDRAVLEVGAAQGVAVGKPSAGGG
GGGATVDAAALAEFTSAITGPDGALAASAHTLLRKLGLETAEGALPDLDDESEKLAALIA
TELGDDWVSAVAPAFDSNKAVLLDDRWASAREDLLRVWNMDPATFPTYSKHILETAKGAG
KALQDQAQWWEKKATEAGRTEHARAYAFLSGLAKSTPEPGVYADEVAVVTGASAGSIAAA
VAGRLLAGGATVICTTSRLDEKRLNFFKELYRTSARTDAALWVVPANMSSYADVDALVEW
IGERQAETVGPYTMVHKEALVPTLLFPFAAPRVSGDMSDAGPRAELEMKILLWSVERLIA
GLSAIGKDTDLASKLHVVLPGSPNRGMFGGDGAYGEAKASLDALVTRWNAEKSWSERVTL
AHAIIGWVRGTGLMGGNDPLVEAVEAAGVRTWSTFEMADQLLENASAEARVRAEQAPLFA
DLTGGLGGVQLDMKALADQAMEARLAEAAEADDEGTIAALPGAPVATLTEPEWDAIDARP
EDMVVIVGAGEVGPYGSSRTRFEMEVDEHLSAAGVAELAWVTGMITWENDPEPGWYDAES
GELVPEAELADKYHDAVVERCGIREFSDDGDIVDTTAPLLVSVFLPEDMTFVVANEAEAR
AFVDADGENTVAVPMETGEWQVTRKAGTEIRVPRRTKLLRSVGGQVPDGFDPSHWGISPE
MLESVDRLAIWNLVATVDAFISAGFEPAELMQYVHPANVANTQGTGMGGMSSMRDLFINT
LLGEPNQNDILQEALPNVVAAHVVQSYVGSYGAMIHPVAACATAAVSVEEGVDKIKVGKA
LFAVTGGFDDLSIEGITGFGMMQATADSEKMLAKGIDPRRFSRANDRRRSGFVESQGGGT
ILLARGDVAAQMGLPVLGVVAWAQSFGDGVHTSIPAPGMGALSVAAGGPQSPMARSLRQL
GVAADDVSIISKHDTSTNANDPNEANLHERIAASIGRSEGAPMFVVSQKSLTGHAKGGAA
AFQLIGMCQMLNDGVVPPNRSLDCVDDEMRKYPHLVWLRETARFGHGFRPKAGLLTSLGF
GHVSGLIAVVHRDAFLAALAPEARSAYLAAAREREVAGRRAVLSAMCGRSELYRKPPAGR
RFGADSSSETTDTREAALLLDDDLRLGQGGTYDGTSCAGGCQ
NT seq
9309 nt
NT seq
+upstream
nt +downstream
nt
atgactgtcgacgacttcgtcaccgcccacgcccgcaccgccaagaacccccgttcgaac
gatgcgctcgtggacaggctggcgtccggcgagcagtacgcgctgatcttcggcggccag
ggcggcccgtggctcgccggtctcgtctcgctgcttgagcagaccggcctggcggacgat
ccggagtaccgcgagctcttcgatggcttcgtctcgcgcgctgacagcgcactcgaaccg
gtgcgggccgagctcttgcgcgcccaagccctcaccttcgacccgatgcggtggcgcgac
gcctcgatcgccgcctccgaggccgagaccaccgacgaggagggcgacgagcccgcccag
gctgccactcaatcgcgccggccccttgccccgcaggtcgacggtgccgtcttctcggtc
cccggcgtcctgttcagccagctcgtggcgctgtacggcctcgaggcccagggcctcgat
ccgtgggacgttccgccggtcgccgccatcggccactcgcagggccagctcgccgtcgcc
gcgttcgcctcccgtggagccgaaggcgacctgctcgccatcgctcgactgatcggtgcc
gccggccgtgtcgtcgcccggcgcaagggcctcgtcgcgctcggcggtgcgaccccgatg
atgtcggtgtccggcgtccgtcaggacaagctcgagcaactcctcgccgacttcaccgct
accgtcgctttcgacgcctgcggccccgtgctcgcggtccgtaatagtcgcaccgcgttc
gtgctcaccggcacccccaagctgctcaacggtttccgcgagtactgcgagtccgtcgcc
gagaccgaggaggcgcagcgcaaggacaagctcaccggcggcgcggcgttcgcccccaag
ttcgaggacgtcccggtcgacatcggtttccaccatcccgcgctcgccgatggcgtcgcg
ctggtcaccgagtgggccgagcgctgcggcctcgacgccgagaaggccggccgcctggcc
gccgccgtgctcgtcgagccggtcgactgggtcgccgatgtcgatgccgcgatcgccaag
ggcgcgcgctggctgatcgatctgggccccgaggatctggcgacccgtctcgccgcggcg
cccgcacgtggccgcggcgtgggactggtgcccgcgaccaccctgcgaggcgcacggaac
ctgttctcccccggaggtattcctgaacttccgctggcctggtcggagttcgggccgcgg
ctggtcgcgttgcccgacggccgcaccgtgctctccacccggttcaccgagctcaccggc
cgcagcccgatcctgctcgccggaatgaccccgaccaccgtcgatccgggcatcgtcgcc
gcagccgcgaacgctggccactgggccgaactcgccggcggcggtcaggtcaccgagccc
atcttcgccgacaacatcgccaagctcaccgcgttgctcgagcccggccgcgctgcgcag
ttcaactcgctgttcctcgacccttacctgtggaagctccagctcggtggcaagcgcatc
gtgcagaaggcccgcgccgcgggcgccccgctcgacggcgtgatcatcaccgcaggcatc
ccggagctcgaggaggccactgccctcgtcgaggagctcaccgacgcgggcatgcggtac
gtcgccttcaagcccggcactgtcgagcagatccgccaggtcgcacggatcgccgccgag
gttccccgctacccggtgatcgtgcacgtcgagggtggaaaggccggtggtcaccactcc
tgggaagacctcgacgaactcctgctctcgacgtacgccgagctgcgtgaccgtggcaat
gtggtgctggccgtgggaggcggcatcggtacccccgaactcgctgccgagtacctgtcc
ggcgaatggtcgcgcaagtacggcgccccggcgatgccggtcgacggcatcctcgtcggc
accgcggcgatggccaccctggaagccgccacctccgacgacgtcaagcggatgctcgcc
gagatccccggcacccccgactggatcggggcgggtcacgcgggcggtggcatggcgtcc
ggccgcagccagctcggtgccgacatccacgagatcgacaacaccgcctccgcatgcggc
cgcctcctcgacgaggtggccggcgatggagacgccgtggccgagcgccgcgacgagatc
gtcgccgccatggccggtaccgccaagccgttcttcggcgacgtcgacgccatgacctac
gcccagtggctcgagcgctacctcgaactctccgtgggcacccgcgcccaggtggccgcc
gccgacccggagcggttcggcgactacggctggctcgacatcacccatcaggaccggttc
ctctcgatgctgcaccgcgccgaggcccgcctcgccgctgaggaccggggcccgatcgag
accgcgttcgccgatgtcaccgccaccgacgatgcctacgccgcgttcgacgctctgttg
cagcgttatccggacgccgcgtccacccgcctgcacccggccgacgtcccgttcttcgtc
tcgctgtgccgcacccccggaaagccggtcaacttcgtgccggtgatcgacaaggatgtg
cgccggtggtggcgttcggactcactgtggcaggcgcactcgcctcgctacaccgccgac
gaggtgtgcatcatccccggcccggaggccgtcgccggaatcacccgcgtcgacgagccg
gtcggcgaactgctgaaccggttcgaggccgaactggtgcgccggcgcacgaacgcgggg
gagcgcgccgaggccgtcgccggtcgccgtgtggtgcccggtgtcgagtccgtgccgctc
gcccgagccctcgccgccaccaacgtgctgtgggccggacggctcgtgcacagccccatc
gcgcgactgggtgcgggcagcacctgggcgatccacggtgagggggacgagtccgtcgcc
ctgcagcccgacaccggcgcccgggtccgcgtgaccggcccgaactcggtgcgcctcgag
attccggtctcgggtaccgaattggtgatcccgatcgaggttccggacagctgcgccgat
ggcggtgtccccgtcatcgagctcggcgaggccagcgaggccatgtccgcactcgtcgca
ttggccgccgccggcgatctgccggcagtccacgacggcgctgccaccatcgatttcgac
ttccgcgcggaggacgccgccgatcacgcggccatcaccgcggacagtctccccgtgggg
ctgtccgccctctacggagacgagcagctggccgtgccggacgccttcgtcggcctgtgc
tggccggcgtcgttcgccgtggtcggcgccgcgaagaccgcagagggacgccccgtcgtc
gagggcatgctggacctggttcacctcgatcacgccatcgagctggccgccaaccccgcg
ccgggctcgtaccgcgccaccgccacgctcgaagacgtgaccgccaccgacatcggcacc
gtggtcaccgtcggcatcgatgtgcgcgcagccggcaccgatgcccccgcggtcaccatg
accgagcggttcgcgatccgtggccgcaccggcgaggcagagctggccgacccgcagccc
gccggcggtgcccgcggtgagaccgccaccgagacgccgcgcaagcgggtccggcaggtc
accgtgcaggccccgcaggacatgagcggcttcgccgcagtctccggcgatcacaacccg
attcacaccagccaggtcgccgcgtcgctcgcaggcctgcccggcccgatcgtgcacggc
atgtggctctcggccgccgcgcagcaggtcacgtccgcctccaacggcgatgtgatgcac
cgtaaacttcgcggctggaccacccgtttcctgggcatggtgctccccggcgacgaggtg
cagttcaccgtcgagcgcgtgggtatcgaccgcggcggcgagctgctcgagatcagcgcc
cgcgtgggcgagaacctggtgatggccgcctccgccgtggccttcccggccgtcaccgcc
tacgtcttccccggccagggcattcagggcaagggtatgggcttggcccagcgctcgcgc
agcaaggccgcccgcgaggtctgggacgccgccgacgagcacacccgcgagaatctcggc
ttctccatcctcgccgtggtccgcgacaatccgctgtcgattcgggccaacggcaccgtc
tacacccatcccgacggggtgctgtacctcaccgagttcacccaggtcgcgatggccacc
gtggcctgtgcacagctggccgaactcaccgagatcggtgcccgcgtggatgatccgatc
atcgcaggccactcggtgggtgagtacaacgcgctcgcgtcggccggggtactgccgctg
ggcaccatcctggagaccgtgttccaccgcggccgcatcatgcagcagctggtgccccgc
gatgcggagggacgcagcccgtacgccatggccgcgatccggcccgagctgttcgaggtc
gacgacgcagacctcgcgaactgggtcgaaggcagcctcgaaggcgcggacgagtacgcc
gagatcgtgaacctgaacctgcgcggcgcgcagtacgtggtggtcggcaatcaggccgga
ctcgacgcattggaggcggcgatcgccgaacgcaccggcagtcccaaggctttccagttg
gtgcccggcatcgatatcccgttccactcgtcggtgctgcgtgccggtgtccccgagttc
gccgcgcgactcgaagagctggtgcccgccgagttcgactacacgaagctggtcggcaag
taccttccgaacctggtggcccgcccgttcgagctgaccgccgatttcgcgctgtcgatc
ctcgacgtggttcccagcgagccggtacgcgagctcgtcgcggattgggagcgcgccgcc
gcgaatccgatggcgaccgcccgcactctgctcatcgagctcctggcctggcagttcgcc
agcccggtccggtggatcgagacccaggatcagctgctgatcgacaccgatcacggcgga
ctgggcgtcgagcggttcatcgaggtggggctcgcgtcggctccgacgctcgcgaacctc
gcgagccgcaccctggcgctggcctcgtatcagcgggccgccgccaccgtgctcaacgcc
gagcgcgatcaggcgaaggtcttcgcgaccgatgaggacgcccccgaggacgaggcggac
gatgccggcaccgaagaggcctcggccgcaccggtttccgtcgccgcggtggcaccgtcg
gccgcgcccgcggcaccggccggtgggccccggcccgacgatctgtccttcgatgcctcc
gacgccactcgcgcgatgatcgcgatctggaccaagatgcgtccggaccagctggagaag
accgacaccatcgagacgctgtgcgacggcgtgagctcgcgccgcaaccagctcctgctc
gacctgggctcggagctcaatctcggtgccatcgacggtgccgcggaggccgatctgccc
acgctggccggcaccgtcaccaaactggcccgcacctacaagccgctcggcccggtgctc
agcgagaccgtcggcaaccagctccgcaagctgctcggttccaccggtaagcgcccgaac
tacatcaccgagtacgtgaccggcacctgggagctcggccccggttgggcgcagcacacc
gtgctcgcactcgccaccggcacccgcgagggcgcttcggtgcgcggcggcgatctggcc
accctgctcgatggcccggtgtccaacgcggctgcgctcgatgcactgatcgaccgtgcc
gtgctcgaagtgggggccgcccagggtgttgccgtcggcaagccctccgccggtggcgga
ggtggcggcgccaccgtcgacgccgcggcgctcgccgagttcacctcggcgatcaccggt
cccgacggtgcgctcgccgcatcggcccatacgctgctgcgcaagctcggcctggaaacg
gccgagggggcattgcccgacctcgacgacgagtcggagaagttggccgcgctgatcgcc
accgaactcggtgacgactgggtcagtgcggtggcgccggccttcgacagcaacaaggcg
gtgctcctggacgaccgctgggccagcgcccgcgaggatctgctccgggtgtggaacatg
gacccggccaccttccccacctacagcaagcacatcctggaaaccgccaagggcgccggc
aaggcattgcaggatcaagcccagtggtgggagaagaaggccaccgaggcgggccgcacg
gagcacgcccgcgcttacgccttcctgtcggggctcgcgaagagcacaccggagccgggc
gtctacgccgatgaggtggccgtggtcacgggcgcctcggcgggatcgatcgcggccgcc
gtcgccggccggctgctggccggcggtgcgaccgtcatctgcaccacgtcgcggctggac
gagaagcggctgaacttcttcaaggagctctaccgcaccagcgcccgtaccgacgcggcg
ctgtgggtggttcccgccaacatgtcgagctacgccgatgtcgacgcattggtcgagtgg
atcggcgagcggcaggccgagaccgtcggcccgtacacgatggtgcataaggaggccctg
gttccgacgctcctgttcccgttcgccgcgccgcgcgtctccggggacatgtccgatgcc
ggcccgcgcgcggagctggagatgaagatcctgctgtggtcggtggagcggctcatcgcc
ggactgtccgcgatcggcaaggacaccgacctcgcgtcgaagctgcacgtggtgctgccc
ggttcaccgaaccgcggcatgttcggcggcgacggcgcatacggcgaggccaaggcgtcc
ctagacgcgctggtcacccgctggaacgcggagaagtcctggtcggagcgggttaccctg
gcgcacgccattatcggatgggtccggggcacgggcctgatgggtggcaacgatccgctg
gtcgaggccgtcgaggccgcaggtgtgcgcacctggtcgaccttcgagatggccgatcag
ctgctcgagaacgcctccgccgaggcccgcgtgcgcgccgagcaggcgccgctgttcgcg
gatctgaccggcggcctgggtggcgtgcagctggacatgaaggccctcgccgaccaggcc
atggaggcccggctcgccgaggccgccgaggcggacgacgagggcaccatcgcggcgctc
ccgggcgccccggtcgccaccctcaccgagcccgagtgggacgccatcgacgcccgcccc
gaggacatggtggtcatcgtcggcgccggcgaggtcggtccgtacggcagttcacgcacc
cggttcgagatggaggtcgacgagcacctctcggccgcgggcgtcgcggagttggcctgg
gtcaccggcatgatcacctgggagaacgatccggagcccgggtggtacgacgccgagtcg
ggcgaactggtgcccgaggcggagctggccgacaaataccacgatgcggtcgtggagcgc
tgcggcatccgcgagttcagcgacgatggcgacatcgtcgacaccacggcaccgctgctc
gtgtcggtcttcctgcccgaggacatgaccttcgtggtcgccaacgaggccgaggcacgc
gccttcgtcgacgccgacggtgagaacaccgtcgccgtgccgatggagaccggcgagtgg
caggtcactcgcaaggcgggcaccgagatccgcgtgccgcgccgcaccaagctgctgcgc
agcgtcggtggccaggtgcccgacggcttcgacccgagccactggggcatctccccggag
atgctggaatcggtcgatcgcctcgcgatctggaatctggtggccaccgtcgacgcgttc
atctccgcgggcttcgaaccggctgagctgatgcagtacgtgcacccggcgaacgtcgcg
aacacccagggcaccggcatgggcggtatgagctcgatgcgcgacctgttcatcaacacg
ctgctgggtgagccgaaccagaacgacatcctgcaggaggcgctgcccaacgtggtcgcc
gcgcacgtggtgcagagctacgtgggcagctacggcgcgatgatccacccggtcgccgcg
tgcgccaccgcggccgtctcggtcgaggagggcgtcgacaagatcaaggtcggtaaggcc
ttgttcgcggtcaccggcggcttcgacgatctgtcgatcgagggcatcaccggtttcggc
atgatgcaggcgaccgccgacagcgagaagatgctcgccaagggcatcgacccgcgccgc
ttctcccgtgccaacgaccggcgccggtcgggcttcgtcgagtcccagggcggcggcacc
atcctgctggcccgcggcgacgtcgccgcccagatggggcttccggtgctcggcgtggtc
gcctgggctcagagcttcggcgacggcgtgcacacctccatcccggcgcccggtatgggt
gcgttgtcggtggcagcgggcggcccgcaatcgccgatggcccgcagccttcggcagctg
ggtgtggcggccgatgacgtgtcgatcatcagcaagcacgacacgtcgaccaacgccaac
gacccgaacgaggcgaatctgcatgagcgcatcgccgcctcgatcggtcgctccgagggc
gcgccgatgttcgtggtctcgcagaagtcgctgaccggccacgccaagggcggtgccgcc
gccttccagctgatcggcatgtgccagatgctcaacgacggtgtggtcccgccgaaccgc
agcctggactgcgtggacgacgagatgcgcaagtacccgcacctggtgtggctgcgcgag
acggcgcggttcgggcacggattccgcccgaaggccggcttgctcacctcgctcggcttc
ggccacgtgtccgggctcatcgcggtggtccaccgcgacgcgttcctggccgcgctcgcc
ccggaggcgcgatcggcgtacctcgcggcggcgcgcgagcgtgaggtggccgggcggcgc
gcggtcctgtcggcgatgtgcggccggtcggagctgtaccgtaagccgcccgccggacgg
cggttcggtgccgattcctcttcggagacgacggacacccgcgaggcggcactgctcctc
gatgacgacctgcgcctggggcagggtggtacgtacgacgggacgtcgtgcgccggtgga
tgccaatag
DBGET
integrated database retrieval system
