Vanessa cardui (painted lady): 124531657
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Entry
124531657 CDS
T09161
Name
(RefSeq) protein dachsous
KO
K16507
protocadherin-16/23
Organism
vcd
Vanessa cardui (painted lady)
Pathway
vcd04391
Hippo signaling pathway - fly
vcd04392
Hippo signaling pathway - multiple species
Brite
KEGG Orthology (KO) [BR:
vcd00001
]
09130 Environmental Information Processing
09132 Signal transduction
04391 Hippo signaling pathway - fly
124531657
04392 Hippo signaling pathway - multiple species
124531657
09180 Brite Hierarchies
09183 Protein families: signaling and cellular processes
04515 Cell adhesion molecules [BR:
vcd04515
]
124531657
Cell adhesion molecules [BR:
vcd04515
]
Cadherins
Cadherin related
124531657
BRITE hierarchy
SSDB
Ortholog
Paralog
Gene cluster
GFIT
Motif
Pfam:
Cadherin
Cadherin_CELSR2_9th
Cadherin_2
RET_CLD1
Cadherin_pro
Cadherin_4
CADH_Y-type_LIR
Motif
Other DBs
NCBI-GeneID:
124531657
NCBI-ProteinID:
XP_046962092
LinkDB
All DBs
Position
8:complement(13633703..13930944)
Genome browser
AA seq
3333 aa
AA seq
DB search
MLRAIICLLVASAAAEHVREFTVRENARAGTLVGHLGDELPDAAPPYTIVPAPGSAVNDD
LKIDPHTGEIRTRVPLDRENRDHYALVAIPASGDNVRVVVRVSDENDNAPTFPAALMNVE
FSENTPRDVKRKLNPAKDEDLGVFNTQRYNIVSGNTDNAFRLSSHRERDGVLYLDLQING
FLDRETTDHYELVIEALDGGTPPLRGTMTVNITILDVNDNPPVFAESAYSAMIPENATVG
TAVLKVFAIDSDAGENGLIEYSINRRQSDKENMFKIDPNTGEITVNKALDFETKELHELV
VVARDKGAQPLETTAFVSIRVTDVNDNQPTIDVIFLSDDATPKISESAQLDEFVARISVH
DPDSKTEYANVNVTLSGGDGHFDLRTHDNIIYLVVVALPLDRESQSSYTLNVVATDKGSP
PLHASRIISLRVTDVNDNPPVFLESVYKTSVPEAAAPGTPVIQVSASDVDEGENSDIRYS
LLPTPQSDWFSIDERSGLVTTRARVDCETNPMPRLTVVASDRGNPPLSSTATLLVTVLDV
NDNEPIFDQSFYNVTVPENEAVGSCILKMSAIDPDCGVNAIVNYTLGDGLSQAFKVRPDT
GELCVSGALDHETNSDFEFPVIATDRGGLSTTAMVKIQLLDINDNVPAFTVKDYDVSLKE
GRISSNEPIVAVSASDADSGRFGTITYRIVNGNDNDVFRIDRSTGEIFVTKPALIQANGK
FDLDVSATDGGGLAAPQAATVHVNVMPAGVGSALFDKPRYNFRVREDVRSGTIVGNLKAT
TGGKQLVRYSIVSGDVDNRFTIEPHTGAIRTSGQLDRETTSSYLLNVKAAVGNPASYDQT
QVQITLEDVNDNAPEFGTSSVRVSVAESAAIGSVVYAARATDEDGGKNGQITYSLVSATG
PSNTFAVNPQHGLVTLLRSLDYENLVRHTLIISAKDNGSPQLSANLTLAVDVQDVNDNAP
IFEHDTYSTNVLESDAVNSKILEIQAIDKDTGNNARITYRVVSENNTNEEHFKVQPTTGW
VYLAKPLDRETIAKHKMTIIATDNGIPPLSATASLIVNVVDANDNDPVFSQTAYEFRVEE
NQKAGAFVGKISATDADLDDNAAIRFSLFPSNTSFVVNPNTGIITTKEILDREFKNSYSL
FAEARDQGSLPRSSRVPISIKITDVNDNSPEIIDPREDVVSVREEQQPGAEVARVKAIDR
DNGLNASVIYSILNERDMDGFGVFVIDSNTGVIKTSTVLDHEERSIYRLTVAATDGGVPP
RQTVRQLKVEVLDLNDNRPTFTSSSLSFKVKEDARIGHIVGVVACCDSDIHENTISDDEE
RQISYLLMSLTSDYSPGTFEIDRRSGSLVVARQLDREIQDEYRLEVRALDTSATNNPQSS
AVTVRIEIIDVNDNAPQWDQDPINIEVSEIISVGTTIYNFTARDEDSGPNGDLNYHITSS
LPVNKNVFNLDPLTGSLSLISPLDYESLKEYWLVVEAVDLATNISERLVTSATVHISVLD
ANDNTPTFVSANKVSVTSNTLSGILYQALAVDTDSGDNGRISYFISGGNDHAYFSIEYNT
GKLTLAKKFSSDLTRVRPGQYKLNLTASDHGIPFPRQSHMTLQLSLQESTNVPPRFTDSY
YRANISEDIRPGSFVTRLSAKSSRGNSGNLSFMIPSGVAGDKFVIDERLGTVTVTTKIDR
EEADQYTFPVYVTDSSTFQSTTNFDVATVSISILDVNDNPPSFKPGSCYTLTVPENNEPE
VIHTVAATDKDIGANGIITYSIISGNNGNKFSIDATTGKLTAKTLDRETQSRYDLTISAY
DHGSPVALQGSCNLTIMVEDQNDNDPVFDTGHYSAAIPEDAAIDTSIIKVRATDADLGYN
KRIVYSLANESQGLFRIDNKTGVIFTTGYFDREKTSSYRFDAVATDQGRYVARSQRVSVE
VTINDVNDNKPIFTKYPFKEQMATLTPPGQSLIRVSASDNDIGINAEILFELLDSSNHKF
RINPTTGVLTATQSLASENGKLIHLKVIAKDKGNPPQSSIGLIELRVGDFSDQTPTLNFQ
NSTYNVTIDENLQYGKDVLQVTALRSDGRRQRIIYEFGNGNDKYAFEIDSNTGVIRVNNS
ANLDYESYPGPRRELVVVARTEGAPILYGYCNVIVNLLDQNDHAPRFTQQQYIANVLEGN
TKGEFVVQLAAKDGDRGANARILYHIVDGNHDNAFIIEPAFSGSVKTNIVLDREIREAYK
LTVIATDEGNPQMTGTATLRINVVDVNDNQPTFPPPNVISVSEGAEVGSVLTSVTANDVD
TYPALKYAIVQGDNTFSIDRYSGKIVLNRPLDYESKKVYEVNISASDSEHVAKTTLTIKV
TDINDNTPVFDDISYNRVLPEGSGTIEIGAVNAKDIDNGDNGKISYSIVQPSPGYYIDSD
TGVLYVNYSALPPHQSDVQLAVIATDHGKPNRSAVASVRINTGASSEIKPFIGQDTYRIT
VNEDTEKGSSLLQIGGINDVLKRYNLDFHIISGNEGDAFDVSTEGALILVNRLDREVQDS
YVLGLAAVEQGKVLSYNQNKTSIMVFVTVTDANDNAPIFSVNDLELTISEDAKIGHTLTQ
LTVTDADLYGTPNSAIIFNITSGDDEKSFYVHPTTGILTVNKSLDYDTGKTKHKLIIVAC
DQGTPCLCNVVYIDIEILDENDNIPTFPVSEYFETIGENERVGTSVFTARATDLDRGKYG
KLNYSIVPLSTNYNRNDDSWKMFQIDSTTGLVNTNAVFDYEQKNKYEFSIKSSDLGGKSS
TVKVRIDIESRDEFYPQFTQKTYNFGVPKSGSLPAGYIIGQATATDRDKGIDGRIVYQLS
TSHSYFKINRTTGVIILKKKVESVKNLFGSDKSISLVVTASSGRQGSLTNKTAVEILLDP
LAVVSDHTNQINDTSIAQSSGLSDWALGLLIAFIFIIIIFAAAFLFLHMKNKRHKKVNKP
GLSSEGVGASNSYVDPSAFDTIPIRNTGPVNSANQFAPPKYDEIPPYGAHTASSNSGAAT
TSELSGSDQSGSSGRGSAEDGEDGEDEEIRMINEGPLQREGLHRQAGGVDDDNLSDVSVH
NTQEYLARLGIVDTGTGGGASTSSRRCSENVGNKDTMLHHGPIDSMHIFDEDGGHENDIT
NLIYAKLNEVTGSERASSADEASAAVDRAMALGAFPSAAGDNTAVPTAGPSMTGSLSSIV
HSEEELTGSYNWDYLLDWGPQYQPLAHVFSEIARLKDDAVSLQSGNSGASSAKSKGTSIS
GGKSVPPPLLTTVAPRSCPAPSLSCRQPQHLLPRSPISHDVPGGFSAAAAMSPSFSPSLS
PLATRSPSMSPLVGPGLPPAPGSRKPPHSTMRM
NT seq
10002 nt
NT seq
+upstream
nt +downstream
nt
atgttgcgcgcaatcatatgcttgctcgtcgcttcggcggcggccgagcacgtcagagaa
tttaccgtccgagagaatgctcgcgccggcaccctcgttggtcacctcggcgacgagctg
ccagacgccgcgccaccatacaccatcgtcccggctccgggtagtgcggtgaatgacgat
ctcaaaatcgatccccacacaggagaaataaggactcgagtgcctttagatagagaaaat
agagaccattacgctctagtcgccatacccgcgagcggtgataacgttcgtgtagtagtc
cgcgtatcggatgaaaacgataacgcgccaaccttccccgccgcactcatgaacgttgag
ttttcggagaatactcctagagacgttaagagaaaactaaatccagccaaagatgaggat
ttaggagtattcaatacacagagatacaatattgtatctggcaacacagataatgcattt
aggttgtcatcgcatagggagcgtgacggcgtgctttatctggaccttcaaatcaatggt
ttccttgatagggaaaccactgatcattatgaattagtcatagaagcgctggatggtggt
acaccaccattaagagggacgatgactgtaaatattaccatattagatgttaacgataat
ccaccggtatttgcggagagtgcatattcagccatgataccagaaaatgctacagttgga
acagctgtgctgaaagtgtttgctatagattctgacgcaggtgaaaatggcttgattgaa
tactcgataaatagacggcagagcgataaagagaacatgttcaaaatagatcctaataca
ggtgaaataactgtgaacaaggctttagactttgagaccaaagaattgcatgagctggtt
gtcgttgctagagataaaggcgctcagcctttggagacgacggcatttgtgtccattcgc
gtcaccgatgttaacgataatcagccaacaattgatgtaatttttctaagtgatgacgct
actcccaaaatatctgaatcagcacagttagatgaattcgtagcgaggatatcagtacac
gatcctgattccaaaactgaatatgcaaacgttaatgttacattgagcggcggtgacgga
catttcgatttacggacgcacgataatatcatatatttagtagtagtggctttacctttg
gacagagagtcccagtcctcctacactttgaatgtcgttgcaacggataagggttcccct
ccattacacgcatctcgcatcatcagtctccgcgtgactgatgttaatgataacccaccg
gtttttctagagagtgtatataaaacaagtgtccctgaagcagcagctccaggcactccc
gttatccaggtgtccgcaagtgatgtagatgaaggggaaaattctgatattaggtattcc
ttacttccaacaccacaatctgattggttttcaattgatgagcgttctggtttggtaacg
acgcgtgctcgtgttgattgtgaaacgaatcccatgccgaggttaacagtggtagccagt
gatcgtggtaatccaccattgtcgtcgaccgcgacgctattggtgacggttttagatgtg
aatgataacgagcctatcttcgaccagtctttctacaacgtgaccgtaccagagaacgaa
gccgtcggcagctgtattttaaagatgtcggcaatcgatccagactgcggtgtgaacgcg
atcgtcaattatacactgggtgacggtctctcccaggcgttcaaagtgagaccggacaca
ggggaactctgtgtgtctggtgctttggaccacgaaacgaacagcgatttcgagtttccc
gtcatagccactgatcgagggggcctgtcgacgaccgcaatggtgaagattcaacttctc
gatataaacgacaacgttccggccttcactgtgaaagattacgacgtttctctgaaggag
ggtaggatatcaagcaacgagccgatagtagccgtgtccgctagcgacgctgattcaggg
agattcggtactataacgtacaggatcgtgaatgggaatgataatgacgtgttccgtata
gataggagtacgggggagatattcgttacgaagccagctctgatacaagctaatggaaag
tttgatttggatgtatcagcgactgatggcggtggtctggccgcgccgcaagcggctacg
gtccatgtgaacgtgatgcctgctggggttggttcggcattgttcgataaaccaagatat
aattttcgtgtacgagaagacgtgaggagcggaaccattgttgggaatctgaaggctacg
accggtggtaagcagctagttcgctacagtatcgtttctggagatgtggataaccgcttc
acaattgagcctcacactggtgcaatcaggacctcggggcaattggacagagagactact
tcttcatatctccttaatgtcaaagctgctgttggcaatccagcgagctatgatcagact
caggtccaaataacattagaagatgtgaacgacaacgctccagaattcgggacgtcttca
gtaagagtatctgtcgctgaatcagcagctatcggttccgtggtgtatgctgcaagagca
actgatgaggatggtgggaaaaatggtcagataacatacagtctcgtctctgctactggc
ccgtcgaacaccttcgctgtgaaccctcaacacggattagtcaccttgttaagatcatta
gattatgagaatctcgttcgtcatacgctcataatatctgccaaagataatggaagcccg
cagttgagtgctaatctaacacttgctgtagatgtccaagatgttaatgataatgctcca
atatttgaacacgacacttacagcaccaatgtgttggaatcagatgctgttaattcaaag
attttagaaatacaagccatagacaaggacaccggtaacaatgcgcgcataacctatcgc
gtcgtatctgagaataatacaaacgaagagcattttaaagttcaaccgactacaggctgg
gtgtacctcgctaaacctctcgatagagagacgatagctaaacacaaaatgacaataata
gctacagacaatggtatcccgccactgtcagcgactgccagccttattgtcaacgtggta
gatgctaatgataatgatccagttttctcacaaaccgcctatgagtttcgagttgaagag
aatcagaaggctggtgcgttcgttgggaagatatcagcgactgatgctgatttggatgat
aatgctgccattagatttagcttgttcccctccaatacgagctttgtcgtcaaccccaat
acaggaattataacaacaaaagaaattttagacagagagttcaaaaattcctattcgctt
ttcgctgaagcgagagatcaaggatccctaccgagatcgagcagagtaccaatatcaatt
aaaataacggacgttaacgacaattcaccagaaataattgatccaagggaagatgttgta
agcgttagagaagaacaacagcctggagcagaagtagctagagtcaaagctattgataga
gacaatggactaaatgcctctgtaatatattcaatcttaaatgagagagatatggatgga
tttggtgttttcgtaattgattcaaatactggcgttatcaaaacgagtacagttttggat
catgaagagagatcaatttatcgtctaacagtcgcagctacagatggaggagtaccacca
agacaaactgtccgacaacttaaagtagaagtactagatttaaatgataatagaccaaca
tttacaagttctagtttatcattcaaagttaaagaagatgccagaattggacatatagtc
ggtgtagtagcatgttgcgatagcgacatacatgaaaatacgataagtgacgacgaggag
agacaaatttcctatttacttatgtcacttacgtcagactattcccctggtacatttgaa
atagacagaagaagtggttctttagttgttgcaagacaattagatagagaaatacaagac
gagtacaggttggaagtgagagccttagatacatcagctacgaataacccgcaaagcagt
gctgtcactgttcgtatcgaaattatcgatgttaacgataacgcaccgcaatgggaccaa
gatcctatcaacattgaggtttctgaaataatatctgttggcacaactatttacaatttc
actgctcgagatgaagattcaggccctaacggagatttgaattatcacataacatcgtcg
ctaccagttaataaaaatgttttcaatttagacccattgaccggttctctgagtctcatt
tcacctctcgattatgagtcattaaaagaatattggttagttgtcgaagccgttgatctt
gcgactaatatatcggagagattggtaacttcagcaacagttcatatttctgttcttgat
gcgaatgataatactcccacttttgtgtcagcgaataaagtatcagtcacatcaaatact
ctttctggtattttataccaagctttggcagttgatacagattcaggagataacggaaga
atatcatatttcatatcaggaggaaatgaccacgcctacttctcaatagaatataatact
ggtaaactaactttagctaagaaattctcttcagaccttacaagagtaagaccaggacaa
tataagctgaatttaactgcttctgatcatggaataccgtttcccagacaaagtcatatg
acattacaattaagcttacaagagtctactaatgtgccaccaagatttacagactcctat
tacagagctaatattagtgaagatattagaccaggtagtttcgtaacgagactatccgct
aaatcatctagaggaaattcaggtaacctgagctttatgataccatctggcgttgctggt
gataaatttgtcatcgatgaacgacttggcacagttactgttaccactaaaattgataga
gaagaagcagatcagtatacttttcctgtttacgttactgattcaagtacttttcagtcc
acgacaaacttcgatgtggctactgtatctataagtattttagacgttaatgacaaccca
ccttcatttaaaccaggctcctgttataccttaacggtacctgagaataatgagcccgaa
gtcatacatacagttgcagcaacagacaaagatataggagcgaatggtataatcacatac
agcattatatctggtaataatggaaacaaattctcgattgatgccacaacgggtaaacta
acagcaaaaactttagacagagaaactcagtcgagatatgacttgactatatctgcgtac
gatcacggttcaccagtggcattacaaggatcatgcaatttaactataatggttgaagac
cagaacgacaacgaccctgtatttgatacaggacattactctgctgcgattcctgaggat
gcagccatagacacgtccattattaaagttagagctactgatgcagatttaggctataat
aaacgaattgtatattcgttagcaaatgaaagtcaaggactatttagaatagataacaag
accggcgttatatttacaactggatactttgaccgtgaaaaaactagtagctatcgtttt
gacgctgttgccaccgatcaaggtcgttatgtagcgcggtcacaaagagtttccgtagaa
gtaacaattaatgacgttaacgacaacaagcctatatttaccaaatatcccttcaaagaa
caaatggctacattaactcctcctgggcaaagtcttatacgtgtttcagccagtgacaat
gacataggcattaacgctgaaatacttttcgaactcttagattcgtcaaatcataaattt
cgcataaacccaactacgggagttcttactgctactcaaagcttggctagtgaaaacggt
aaattaatacacctcaaagtaatagcaaaagacaaaggaaatcccccgcagagctcaata
ggtcttatcgaattacgtgttggagatttttctgatcaaacaccaacccttaactttcaa
aattcgacttacaatgtaacaattgacgaaaatttacaatacggaaaggatgttctacaa
gtaacagcactgagaagtgacggacgtcggcagagaattatttatgaatttggcaatggt
aatgacaagtatgcctttgaaatagactctaataccggcgtcataagggttaacaactca
gcaaatttagattatgaatcatatccgggaccaagaagggaactagtagttgtagccaga
acagaaggcgctcctattttgtatggttattgtaacgtaatagttaaccttttggaccaa
aacgatcacgctcctaggtttacacaacaacaatacatagctaatgttttagaaggaaac
acaaagggcgagtttgttgttcaattagctgctaaagatggagatcgaggagcaaatgcg
cgaatactttatcacatcgttgatggtaatcacgataacgcatttattatagagcctgcc
ttttcgggttcagtaaaaaccaacatagttctcgatagagaaataagagaagcttataaa
ctaacagttatagcgactgatgaaggcaatcctcagatgacggggactgccactctcaga
attaacgtcgtcgatgtcaatgacaatcagccgacattcccaccgccaaatgtcatctca
gtaagcgaaggagcggaagtcggctcagtacttacatctgttactgcaaatgatgtcgac
acgtatccagctctgaaatatgcgattgttcaaggagataacactttttcaatagacaga
tatagtgggaagattgtcttgaacaggccacttgattatgaaagcaaaaaagtatatgaa
gtgaatataagcgcatctgatagtgaacacgttgcgaagacaactttaacaatcaaagtg
acagacattaatgataacacgcccgtttttgatgacatttcatataacagagtcttgcct
gaaggcagcggcactattgaaatcggcgcagtcaacgcaaaagatattgacaatggggat
aacgggaagatttcgtactccattgtgcaaccaagtcctggctactatattgattctgat
accggggttttgtatgtgaactacagcgctttgccccctcaccagagtgatgtgcaatta
gctgtgattgcaacggaccacggcaaaccaaatagaagtgctgtggcatcagtgaggatc
aacactggcgcttcgtcggaaataaaaccttttatcggacaagatacttacagaatcaca
gtcaatgaagacacggagaagggatcatcacttcttcaaattggcggaattaacgacgtt
ctcaagagatacaaccttgacttccacataatatccggaaatgagggcgatgcctttgat
gtcagtacagaaggagccttgatacttgtcaatagattagacagagaggtgcaagattca
tacgtccttggcttagctgcagtagaacaaggcaaggttttgtcttacaatcaaaataaa
acgtcaatcatggtatttgttactgtaacagatgccaatgacaacgctccgatattttct
gtaaatgatttagagctcaccatcagcgaagatgccaaaattggtcatacgctaactcaa
ctaacagtgaccgatgctgatttgtatggtacaccaaactccgcgattatttttaatata
acttcaggagacgatgaaaaatcgttttacgtgcatccaacaactggtattttgactgtt
aataaatcactcgactatgatacaggaaaaacaaaacacaaattaataatagtcgcctgt
gatcaaggcactccgtgcctttgtaacgttgtgtacatagatatcgaaattctcgacgaa
aacgataatatacctacattccccgtcagtgaatatttcgaaacaataggcgagaatgaa
cgcgtcggaacatcggtattcacagcaagagctacagatcttgacagaggcaaatatgga
aaactaaattactctatagtaccattgtctacaaattacaatagaaacgatgactcctgg
aaaatgtttcaaatagattccacaacaggtcttgtgaatactaacgcagtattcgactat
gaacaaaagaacaaatatgagttctctataaagtcatcagatttaggcggtaaaagttct
acggtcaaagtgagaattgacatagaaagtcgagacgagttttatccccagtttactcag
aaaacctacaattttggtgtcccgaaatctggttcattaccggcgggttacattatagga
caagcgacggcaactgaccgagacaaaggtatcgatgggcgcatagtgtatcagctgtca
acttcacattcatactttaagattaatagaacaacaggcgtcataattttgaaaaagaaa
gtagaatcggtcaaaaacctattcggttctgataaatcgataagcttagtggttactgcc
agttcaggcaggcaaggctctcttacgaataaaactgcagttgaaatattacttgatcct
ttagcagttgtctccgaccacacaaatcagatcaacgacacgtctattgcacaaagcagt
ggactttccgattgggcattaggattattaattgcatttatatttattattataatattt
gcagcggcattcttatttttacatatgaaaaataaaaggcataagaaagtcaataagcca
ggactcagttctgaaggagttggagcgtcaaatagttatgtagaccccagtgcttttgat
acaatcccaataagaaatacgggtccagtgaattccgccaatcaatttgctccccctaaa
tatgacgagatacctccatacggggcccatacagcgagctctaattcaggagctgccaca
acttctgaactatctggttctgatcaatctggatcaagtggccgtggatcagcagaagat
ggagaagacggtgaagatgaggaaattagaatgataaacgagggaccgttgcaacgagaa
ggattacatagacaggctggtggagtcgacgatgacaacctttctgatgtttctgttcat
aacactcaagaatatttagctcgattaggaattgtggatactgggaccgggggaggagca
tcaacatcatctcgaagatgttcagaaaatgttggtaataaagacacaatgttgcatcac
ggaccaattgattcaatgcacatatttgacgaagatggcggtcacgaaaacgatataaca
aacttaatttatgctaaacttaatgaagttaccggcagcgagagagctagtagtgctgat
gaagcaagtgctgccgtagatagagcgatggccttaggtgctttcccaagtgcagcaggt
gataatactgctgtacctactgctggtccatctatgacgggaagtctaagctctattgtt
cattcagaagaagaattgactggaagctataactgggattatttactagattggggacca
caatatcaacctttagcacatgttttctctgaaattgcacgtttaaaagatgacgcggtg
tcattacaaagtggtaacagtggtgcatcaagtgctaagagtaaaggaacatcaatatcc
ggtgggaagagtgtccctccgccattactcacaacagttgcaccaagaagttgtcctgct
ccatcattatcatgtagacaaccacagcacttattaccaaggtctcctattagtcatgat
gttcctggtggattttccgcagctgctgctatgtcaccgtccttttcgccatcattatct
ccattagctactcgatcaccatctatgtcgccacttgtagggccaggtcttccgcctgcg
ccgggaagtagaaaacctcctcactccacaatgaggatgtaa
DBGET
integrated database retrieval system