Entry
Name
6-pyruvoyltetrahydropterin synthase;
2-amino-4-oxo-6-[(1S,2R)-1,2-dihydroxy-3-triphosphooxypropyl]-7,8-dihydroxypteridine triphosphate lyase;
6-[(1S,2R)-1,2-dihydroxy-3-triphosphooxypropyl]-7,8-dihydropterin triphosphate-lyase (6-pyruvoyl-5,6,7,8-tetrahydropterin-forming)
Class
Lyases;
Carbon-oxygen lyases;
Acting on phosphates
BRITE hierarchy
Sysname
7,8-dihydroneopterin 3'-triphosphate triphosphate-lyase (6-pyruvoyl-5,6,7,8-tetrahydropterin-forming)
Reaction(IUBMB)
7,8-dihydroneopterin 3'-triphosphate = 6-pyruvoyl-5,6,7,8-tetrahydropterin + triphosphate [RN:
R04286 ]
Reaction(KEGG)
Substrate
7,8-dihydroneopterin 3'-triphosphate [CPD:
C04895 ]
Product
6-pyruvoyl-5,6,7,8-tetrahydropterin [CPD:
C03684 ];
triphosphate [CPD:
C00536 ]
Comment
Catalyses triphosphate elimination and an intramolecular redox reaction in the presence of Mg2+. It has been identified in human liver. This enzyme is involved in the de novo synthesis of tetrahydrobiopterin from GTP, with the other enzymes involved being EC
1.1.1.153 (sepiapterin reductase) and EC
3.5.4.16 (GTP cyclohydrolase I) [3].
History
EC 4.2.3.12 created 1999 as EC 4.6.1.10, transferred 2000 to EC 4.2.3.12, modified 2001
Pathway
Orthology
K01737 6-pyruvoyltetrahydropterin/6-carboxytetrahydropterin synthase
K22101 dihydroneopterin triphosphate aldolase (PTPS-III) / 6-pyruvoyltetrahydropterin synthase
Genes
ANAN : 105720753 105729610(PTS)
DSP : 122097818 122110301(Pts)
NPO : 129496921(PTS) 129508266
NVS : 122889286 122912285(PTS)
ZCA : 113915515(PTS) 113927632
MLX : 118002115 118013239(PTS)
NSU : 110576930 110586387(PTS) 123325711
BTA : 101905029 506860(PTS)
BOM : 102280040 102281235(PTS)
BIU : 109569917(PTS) 109570913
BBUB : 102399912(PTS) 102404326 102410224
BBIS : 104981484(PTS) 104992004
BTAX : 128060838(PTS) 128070603
ODA : 120876258(PTS) 120880189
CCAD : 122441826 122450191(PTS)
MBEZ : 129554336(PTS) 129564214
MJV : 108392590 108401540(PTS)
ASN : 102370397(PTS) 102370641
AMJ : 102570435(PTS) 106737502
CPOO : 109317947(PTS) 109318087
GGN : 109292974(PTS) 109292980
PSS : 102443610 102453541 102453775
CABI : 116824038 116831007
ACS : 100567950 107983749(pts)
ASAO : 132782463 132782670 132782671
PVT : 110078745 110078754 110078762
SUND : 121934418 121934745 121935519
PBI : 103053109(PTS) 103053343
PMUR : 107284508(PTS) 114922340
CTIG : 120311254(PTS) 120311255
PGUT : 117670167 117670238(PTS)
APRI : 131203601(PTS) 131203928
PTEX : 113441817 113441949(PTS)
NSS : 113420850(PTS) 113420902
VKO : 123036843 123036886 123036887
PMUA : 114585329 114585330 114585688
PRAF : 128402574 128402575 128402576
ZVI : 118097078 118097079 118097357
HCG : 128333548 128333621 128334041
STOW : 125441765(PTS) 125441766 125441767
EMC : 129342291 129342327(PTS) 129342328
XLA : 108715196(pts.S) 495178(pts.L)
SRX : 107736111(pts) 107742600
CAUA : 113068189 113112019
CGIB : 127969400 128025740
PLEP : 121939447 121958525(pts)
MSAM : 119915738 119916168(pts)
CVG : 107081163 107095326(pts)
KMR : 108235842(pts) 108236896
SLAL : 111650306(pts) 111652555
CCLU : 121547984(pts) 121581361
RTP : 109912892 125485856(pts)
SKO : 100370945 100376492 100378870
DSI : Dsimw501_GD28180(Dsim_GD28180)
ECOE : 129940829 129942312
AGIF : 122857934 122860081
CCIN : 107269365 107269395
PPYR : 116164520 116172257 116172260 116182628
DPL : KGM_215764 KGM_215765
CFEL : 113367277 113367700 113381749
TPAL : 117644579 117644580
SGRE : 126271848 126323730
BROR : 134532610 134533377
CSCU : 111627504 111627506
CEL : CELE_B0041.6(ptps-1)
CBR : CBG_22232(Cbr-ptps-1)
BMY : BM_BM1701(Bma-ptps-1)
HRF : 124134198 124134200 124134201 124134244 124134260
RPHI : 132718746 132718748 132718750 132718751 132718754 132730226
AMIL : 114972416 114977768
AQU : 100633397 105315318 109582097 109584937
PLAG : 39740896(PADL01_0626200)
PMAL : PMUG01_11035600(PTPS)
PBRS : 92371779(MKS88_003692)
PREL : PRELSG_1120300(PTPS)
EOH : ECO103_3309(ygcM) ECO103_5074
ECOO : ECRM13514_3621(ygcM)
ECOH : ECRM13516_3487(ygcM) ECRM13516_5069
ECW : EcE24377A_3067(queD)
EDJ : ECDH1ME8569_2675(ygcM)
ECOI : ECOPMV1_03018(queD)
ESZ : FEM44_03725(queD) FEM44_14100(queD)
SENC : SEET0819_06395 SEET0819_21580
ECLG : EC036_05180 EC036_35280
ECAN : CWI88_04600(queD) CWI88_19485(queD)
ENX : NI40_002635 NI40_017670
KVQ : SP68_09785 SP68_17575
REE : electrica_00939(queD)
RTG : NCTC13098_01416(queD)
CFQ : C2U38_02650(queD) C2U38_20035(queD)
CFAR : CI104_20010(queD) CI104_21860(queD)
CENS : P2W74_04665(queD) P2W74_20150(queD)
CNT : JT31_15265 JT31_16650
CLAP : NCTC11466_00738(queD)
ESC : Entcl_0941 Entcl_3865
KOT : EH164_17830(queD) EH164_19590(queD)
KIE : NCTC12125_04928(queD)
LAZ : A8A57_02655 A8A57_16735
BAGE : BADSM9389_09140(queD)
AHN : NCTC12129_03960(ygcM)
METY : MRY16398_10330(ygcM)
PTF : PROFFT_A_03080(queD)
PSHI : SAMEA2665130_1260(queD)
PVJ : LMA04_15075(queD) LMA04_16490(queD)
SMW : SMWW4_v1c07910(queD)
SFJ : SAMEA4384070_0840(queD)
SOF : NCTC11214_00694(queD)
SSAR : SSARUM_000734(queD)
SERA : Ser39006_007300(queD)
PAQU : DMB82_0004550(queD)
PARL : PEC302110_31340(queD)
DIC : Dpoa569_000791(queD)
BNG : EH206_04390(queD) EH206_20430(queD)
EHD : ERCIPSTX3056_601(queD)
BAPF : BUMPF009_CDS00182(sscr)
BAPG : BUMPG002_CDS00182(sscr)
BAPU : BUMPUSDA_CDS00182(sscr)
BAPW : BUMPW106_CDS00182(sscr)
TPTY : NCTC11468_00907(queD)
PLUM : A4R40_03510 A4R40_13930
PAKH : B0X70_04025 B0X70_14245
PLUI : CE143_03960(queD) CE143_14240
PHEI : NCTC12003_00941(queD)
PRJ : NCTC6933_03111(queD)
HIC : NTHIC486_01555(queD)
HPIT : NCTC13334_01342(queD)
HAEG : NCTC8502_01316(queD)
PDAG : 4362423_01165(queD)
ADP : NCTC12871_00351(queD)
ALIG : NCTC10568_01371(queD)
AVT : NCTC3438_01934(queD)
RPNE : NCTC8284_02630(queD)
PAET : NCTC13378_01691(queD)
XFU : XFF4834R_chr35090(ptpS)
XPH : XppCFBP6546_16260(queD)
XHD : LMG31886_37030(queD)
LLZ : LYB30171_00447(queD)
TBV : H9L17_06735(queD) H9L17_11170
THEX : ICG51_000985 ICG51_001775(queD)
VAU : VANGNB10_cI1399c(queD)
VPL : SA104470976_01044(queD)
PAEB : NCGM1900_3845(ptpS)
PVD : CFBP1590__3312(queD)
PPRC : PFLCHA0_c40780(queD)
PMUD : NCTC8068_03259(queD)
PFW : PF1751_v1c19380(queD)
PTRT : HU722_0010780(queD)
PTAE : NCTC10697_03086(queD)
PGF : J0G10_17115(queD) J0G10_20145(queD)
PNB : NK667_05670(queD) NK667_16225(queD)
PSOA : PSm6_00500 PSm6_18830
PKJ : Q1W70_00140(queD) Q1W70_01895(queD)
PSJY : AA098_03725 AA098_10070
PBAM : NUH87_27695(queD) NUH87_28365(queD)
AVN : Avin_28600 Avin_29180
AVL : AvCA_28600 AvCA_29180
AVD : AvCA6_28600 AvCA6_29180
PSPI : PS2015_1143 PS2015_1899
SPOI : IMCC21906_02523 IMCC21906_02802
SNAN : I6N98_04470 I6N98_07200(queD)
STAW : NCG89_13495 NCG89_15325(queD)
ZAL : AZF00_04545 AZF00_06850
OSG : BST96_14735 BST96_15940
HJA : BST95_10100 BST95_18015
HALC : EY643_04110 EY643_12515(queD)
KIM : G3T16_14355 G3T16_18605(queD)
AFUS : EYZ66_05000 EYZ66_05235(queD)
PAQM : E0F26_10615(queD) E0F26_11790
MDN : JT25_000475 JT25_015580
MDH : AYM39_07755 AYM39_14935
MKO : MKLM6_1597 MKLM6_3073
METL : U737_05175 U737_10915 U737_15205(queD)
MELL : IVG45_11265(queD) IVG45_13770
MMOT : QZJ86_04895 QZJ86_09450
MAH : MEALZ_0719(ygcM) MEALZ_2440
MBUR : EQU24_03540 EQU24_08610
MSZE : MSZNOR_1589 MSZNOR_2904
MISZ : MishRS11D_33030(queD)
MCAU : MIT9_P0854 MIT9_P1579
ALV : Alvin_1560 Alvin_2762
ATEP : Atep_16960 Atep_28430
TVI : Thivi_0093 Thivi_3266
MPUR : MARPU_08795 MARPU_11395 MARPU_12765
TTP : E6P07_09200 E6P07_11845
THIP : N838_01205 N838_21995
CJAP : GWK36_06715 GWK36_13230
THIM : KFB96_14085 KFB96_25435
TBOG : LT988_02380 LT988_09610
TNI : TVNIR_2462(queD_[H])
THIC : TspCOW1_14700(queD)
NMM : NMBM01240149_1577(queD)
NMS : NMBM01240355_0524(queD)
NMQ : NMBM04240196_1639(queD)
NMZ : NMBNZ0533_0564(queD)
RME : Rmet_1101 Rmet_1572(ptpS)
BPM : BURPS1710b_0358(queD)
BPSE : BDL_1814(queD) BDL_4886
BPSD : BBX_204(queD) BBX_4048
BPK : BBK_1292(queD) BBK_4189
BUR : Bcep18194_A6468 Bcep18194_B0318
BCM : Bcenmc03_3133 Bcenmc03_4938
BCH : Bcen2424_3117 Bcen2424_5330
BCEN : DM39_3216(queD) DM39_5308
BMK : DM80_1793(queD) DM80_4777
BCED : DM42_1894(queD) DM42_5451
BCEP : APZ15_03500 APZ15_23185
BCON : NL30_16455 NL30_28715
BSEM : WJ12_15615 WJ12_32675
BPSL : WS57_01485 WS57_34405
BMEC : WJ16_15805 WJ16_31150
BSTL : BBJ41_13000 BBJ41_26625
BANN : JFN94_15945(queD) JFN94_16980
BARI : NLX30_17825(queD) NLX30_24475
BAEN : L3V59_17745(queD) L3V59_27610
BGF : BC1003_0008 BC1003_0080
BGD : bgla_1g12780 bgla_1g38220
BGO : BM43_1338(queD) BM43_2611
PARA : BTO02_01210 BTO02_22955
PLAD : PPGU16_00390 PPGU16_01600
PPUL : RO07_09275 RO07_10285
PSPU : NA29_05075 NA29_06100
PAPI : SG18_09815 SG18_10700
PVE : UC34_00915 UC34_01965
POX : MB84_05280 MB84_06405
PTX : ABW99_16355 ABW99_16830
PNR : AT302_08955 AT302_10030
PFIB : PI93_000920(queD) PI93_001735
PCOM : NTU39_01295(queD) NTU39_02230
PLG : NCTC10937_03552(queD)
KIA : G8A07_00055 G8A07_00415
JLV : G3257_09360(queD) G3257_16805(queD)
AON : DEH84_00165 DEH84_02900
METR : BSY238_1605 BSY238_1793(queD)
URU : DSM104443_00488(queD)
QAU : KI612_08820 KI612_15955(queD)
APP : CAP2UW1_1018 CAP2UW1_4261
REN : EF513_00745 EF513_02300
MLN : A9174_32275 A9174_32995
MJR : EB229_32995(queD) EB229_33755(queD)
MERD : EB233_30630(queD) EB233_31415(queD)
NRH : T8J41_03895(queD) T8J41_09055
AMIS : Amn_09990 Amn_11270 Amn_pa00870
RBS : RHODOSMS8_00914(queD)
SMEL : SM2011_b20939(exsC)
SFD : USDA257_c04430(queD) USDA257_c13700(exsC)
EAK : EKH55_4941 EKH55_5754
ALF : CFBP5473_17410(queD)
AVV : RvVAT039_10260(exsC)
ASAL : CFBP5507_22295(queD)
RPUS : CFBP5875_21280(queD)
ALEG : CFBP4996_16195(queD)
REC : RHECIAT_CH0003985(exsC)
REL : REMIM1_CH03790(queD)
REP : IE4803_CH04098(queD)
REI : IE4771_CH04039(queD)
RTR : RTCIAT899_PC05460(queD)
RGA : RGR602_PC02215(queD)
RPHA : AMC79_CH03915(queD)
RJG : CCGE525_32605(queD) CCGE525_34805(queD)
RLW : RlegWSM1455_19115(queD)
RBW : RLCC275e_19420(queD)
NEN : NCHU2750_27570(exsC)
BIO : BR141012304_10744(queD)
BEL : BE61_20440 BE61_29940
BSEP : HAP48_0006830 HAP48_0035055 HAP48_0049265
BCAN : BcanWSM471_06575(queD) BcanWSM471_11495
BXN : I3J27_08615 I3J27_33950 I3J27_33960(queD)
PCAX : AFIC_002396 AFIC_002538(queD)
VGO : GJW-30_1_02582(queD)
BOF : FQV39_07765 FQV39_29010
MPHY : MCBMB27_01591 MCBMB27_05609
METX : A3862_21100 A3862_29715
MMES : MMSR116_25490(queD)
MOC : BB934_11630 BB934_43240
MLD : U0023_08860 U0023_25635 U0023_27795 U0023_29090
MTUN : MTUNDRAET4_2149(ygcM)
FIL : BN1229_v1_2819(queD)
FIY : BN1229_v1_3094(queD)
CAUL : KCG34_18930 KCG34_25425
BDM : EQG53_00575 EQG53_01050
BMED : GYM46_02540 GYM46_14225
BVY : NCTC9239_00387 NCTC9239_01366
BND : KWG56_05860 KWG56_08975
BRES : E4341_06325 E4341_12785
PHP : PhaeoP97_02794(queD)
PPIC : PhaeoP14_00483(queD)
SINL : DSM14862_02082(queD)
RMM : ROSMUCSMR3_03728(queD)
LVS : LOKVESSMR4R_00078(queD)
PPRU : FDP22_22520 FDP22_23920 FDP22_24295
RGI : RGI145_04290 RGI145_04305
ROS : CTJ15_13100 CTJ15_13115
RMUC : FOB66_06090 FOB66_06105
SROE : JL101_000535 JL101_029360
HPYE : HPOKI154_00385 HPOKI154_04735
HPYG : HPOKI673_00385 HPOKI673_04720
HPYH : HPOKI828_00385 HPOKI828_04720
CJV : MTVDSCj20_0158(queD)
CFV : CFVI03293_0224(queD)
CAMP : CFT03427_0231(queD)
CCOC : CCON33237_0250(queD)
CLR : UPTC16701_1395(queD)
CLM : UPTC16712_1415(queD)
CAMR : CAQ16704_1432(queD)
CHV : CHELV3228_1502(queD)
CPIN : CPIN18020_1094(queD)
CAMZ : CVIC12175_1385(queD)
AFC : AFAEC_0257(queD) AFAEC_a0014
AAQI : AAQM_0756 AAQM_2220(queD)
ACLO : ACLO_1750 ACLO_2353(queD)
AVP : AVENP_0625 AVENP_2736(queD)
ADZ : ADFLV_2301 ADFLV_2660(queD)
SMAB : LN246_01300 LN246_13840
GEC : GEO60473_11610(queD)
GBN : GEOBRER4_28100(queD)
GPL : M1B72_02670(queD) M1B72_06290(queD)
PEF : A7E78_02175 A7E78_06910
DPG : DESPIGER_1120 DESPIGER_2411
DSB : LN040_00325(queD) LN040_08530
DCB : C3Y92_07625 C3Y92_14235
DVG : Deval_1247 Deval_2449
CLIH : KPS_000151(queD) KPS_001391(queD)
DESU : NLA06_00495(queD) NLA06_05110
DPS : DP0165 DP2265 DP2266
DTO : TOL2_C37050(ptpS) TOL2_C37530(queD)
DML : Dmul_05960(queD) Dmul_37770(queD)
DEK : DSLASN_09490 DSLASN_40800(queD)
DTI : Desti_0247 Desti_0743 Desti_1219 Desti_2993
DAX : FDQ92_04120(queD) FDQ92_09770(queD)
ACP : A2cp1_1437 A2cp1_1997
AFW : Anae109_1354 Anae109_1887
ANK : AnaeK_1339 AnaeK_1912
AORY : AMOR_07690 AMOR_57950
APAU : AMPC_02190(queD) AMPC_05710 AMPC_17350
MSD : MYSTI_05526 MYSTI_05528
MLAN : JY572_39630(queD) JY572_39640
CCX : COCOR_01408 COCOR_01410(queD)
MBD : MEBOL_002464 MEBOL_004769 MEBOL_004771
CFUS : CYFUS_001868 CYFUS_001870
SCL : sce5728(ygcM1) sce8620(ygcM3)
SCU : SCE1572_34050 SCE1572_46590
LLU : AKJ09_02760 AKJ09_03724
NANN : O0S08_32185(queD) O0S08_32190 O0S08_50685(queD)
BBA : Bd1863(ygcM) Bd1864 Bd1968(Pts)
BBAT : Bdt_1832(ygcM) Bdt_1833 Bdt_1939(pts)
BBAC : EP01_05385 EP01_05390 EP01_05885
BDQ : CIK05_09130 CIK05_09135
BDC : DOE51_08830(queD) DOE51_08835 DOE51_09360
BDZ : DOM22_10470 DOM22_11120 DOM22_11125(queD)
BDK : HW988_09775 HW988_10305 HW988_10310(queD)
BREY : MNR06_03990 MNR06_03995(queD)
BAZ : BAMTA208_10270(queD)
BSON : S101395_02366(queD) S101395_03385
BMYC : DJ92_3983(queD) DJ92_772(queD)
BWD : CT694_07200(queD) CT694_32700(queD)
BGY : BGLY_1466(queD) BGLY_2486
BXI : BK049_01730 BK049_08180
BFD : NCTC4823_01601(queD)
BACW : QR42_02370 QR42_06485
BAEI : RE735_00440 RE735_13050(queD)
GTK : GT3570_04715 GT3570_07080
BCAL : CWI35_02825 CWI35_05220(queD)
PARG : PspKH34_02950 PspKH34_28220(queD)
ACAI : ISX45_02150(queD) ISX45_08015
BLEN : NCTC4824_01075(queD)
DOM : RRU94_03180 RRU94_12725(queD)
PUK : PU629_16985(queD) PU629_20615
SUQ : HMPREF0772_12476(queD)
SHH : ShL2_00084 ShL2_02035
SLG : SLGD_00250 SLGD_02123
SLN : SLUG_02490 SLUG_20950
SSD : SPSINT_0420 SPSINT_2129
SDP : NCTC12225_00417 NCTC12225_02255(queD)
SXY : BE24_01305 BE24_10510
SXL : SXYLSMQ121_0317 SXYLSMQ121_1988
SXO : SXYL_00321 SXYL_02140(queD)
SHU : SHYC_02255 SHYC_10170(queD)
SSCH : LH95_02005 LH95_09690
SSCZ : RN70_02320 RN70_10455
SAGQ : EP23_07880 EP23_11980(queD)
SEQO : SE1039_05350 SE1039_22580
SPET : CEP67_08395 CEP67_10495(queD)
SCOH : BZ166_02000 BZ166_06095
SURY : MUA21_02220(queD) MUA21_11165
SNL : BJD96_01900 BJD96_11320
SKL : C7J89_03385 C7J89_11930(queD)
SFQ : C7J90_01190(queD) C7J90_04810
SMUS : C7J88_06335 C7J88_08975(queD)
SCHR : DWB92_02080 DWB92_10325(queD)
SARL : SAP2_04160 SAP2_20640
SPIC : SAMEA4384060_2159(queD)
SSH : NCTC13712_00676(queD)
SSIM : SAMEA4384339_0190 SAMEA4384339_0656(queD)
SLLO : ISP08_02135 ISP08_10200(queD)
SAUL : I6G39_09220(queD) I6G39_10345
SRAI : LN051_01725 LN051_09460(queD)
SPSD : JMB28_02105 JMB28_10845(queD)
SEDA : MNY58_01660 MNY58_10870(queD)
SCAQ : QMO72_01755 QMO72_10760(queD)
SDEV : Q2T90_06435 Q2T90_08785(queD)
STAP : AOB58_1219 AOB58_2144
SGAI : K3U27_01240(queD) K3U27_07885
SFF : FOB90_05760 FOB90_09760(queD)
SSTE : SAMEA4384403_2105(queD)
MACR : BHM04_07255 BHM04_07730
MARY : LAU42_02375 LAU42_02750(queD)
MBRU : MGG13_00600 MGG13_03010(queD)
MEPI : KFV12_03170 KFV12_03680(queD)
MBOC : QSV55_01005 QSV55_02910(queD)
MCAK : MCCS_05890 MCCS_06680(queD)
MBOE : HT586_03070 HT586_03545(queD)
BHUI : LOK74_08260 LOK74_15085(queD)
BAYD : BSPP4475_04110(queD)
PPM : PPSC2_01040(ygcM) PPSC2_05570(ygcM)
PVO : PVOR_07295 PVOR_09220
PCHI : PC41400_05710(queD)
PBK : Back11_39430 Back11_48180(queD)
PPRT : ET464_01280(queD) ET464_04295
PBAC : HUB98_16520 HUB98_22825(queD)
PRZ : GZH47_28000(queD) GZH47_28025(queD)
PNK : AASFL403_13920(queD)
PKP : SK3146_01933(queD_1) SK3146_02868(queD_2)
PGLU : A3958_14085 A3958_15675
ANX : ACH33_11310 ACH33_14235
ATHE : K3F53_04675(queD) K3F53_07695
ANEU : PP175_07035(queD) PP175_25930
AACO : K1I37_03110 K1I37_13615(queD)
EFF : skT53_00110 skT53_15520
SGG : SGGBAA2069_c12880(ptpS)
SVB : NCTC12167_00834(ptpS)
CAE : SMB_G2620 SMB_G3665(queD)
CBA : CLB_0868(queD) CLB_1575(queD)
CBY : CLM_0969(queD) CLM_1795(queD)
CBL : CLK_0229(queD) CLK_1033(queD)
CBB : CLD_2998(queD) CLD_3745(queD)
CBI : CLJ_B0869 CLJ_B1664(queD)
CBF : CLI_0909(queD) CLI_1636
CBM : CBF_0880(queD) CBF_1617
CBJ : H04402_00887 H04402_01621
CLD : CLSPO_c08280(queD) CLSPO_c15970
CALE : FDN13_03665 FDN13_03705(queD)
CALD : PWK10_03510(queD) PWK10_16740(queD)
CEV : LK421_03150(queD) LK421_13225
EEL : EUBELI_20139 EUBELI_20445
VRM : 44547418_01275(queD)
VDN : NCTC11831_00683(queD)