KEGG ORTHOLOGY: K22516

ORTHOLOGY: K22516

Entry

K22516                      KO

Symbol

fdhA

Name

formate dehydrogenase (coenzyme F420) alpha subunit [EC:1.17.98.3 1.8.98.6]

Pathway

map00680

Methane metabolism

map01100

Metabolic pathways

map01120

Microbial metabolism in diverse environments

map01200

Carbon metabolism

Module

M00356

Methanogenesis, methanol => methane

M00357

Methanogenesis, acetate => methane

M00563

Methanogenesis, methylamine/dimethylamine/trimethylamine => methane

M00567

Methanogenesis, CO2 => methane

Reaction

R10797

formate:coenzyme-F420 oxidoreductase

R11944

coenzyme B,coenzyme M,ferredoxin:formate oxidoreductase

Brite

KEGG Orthology (KO) [BR:ko00001]
09100 Metabolism
  09102 Energy metabolism
   00680 Methane metabolism
    K22516  fdhA; formate dehydrogenase (coenzyme F420) alpha subunit
Enzymes [BR:ko01000]
1. Oxidoreductases
  1.8  Acting on a sulfur group of donors
   1.8.98  With other, known, physiological acceptors
    1.8.98.6  formate:CoB-CoM heterodisulfide,ferredoxin reductase
     K22516  fdhA; formate dehydrogenase (coenzyme F420) alpha subunit
  1.17  Acting on CH or CH2 groups
   1.17.98  With other, known, physiological acceptors
    1.17.98.3  formate dehydrogenase (coenzyme F420)
     K22516  fdhA; formate dehydrogenase (coenzyme F420) alpha subunit

Other DBs

COG:

COG3383

GO:

0043794

Genes

VNA:	PN96_21155

VEJ:	VEJY3_20411

VJP:	NP165_14560(fdhF)

VIB:	VspSTUT11_44260

VID:	DBX26_16030

PGB:	H744_1c0419

PGH:	FH974_23235(fdhF)

PHOT:

L4174_007680(fdhF)

PCH:	EY04_14750

PPHN:

HU825_03470(fdhF)

PSH:	Psest_3905

PSTU:

PSED:

SDEG:

GNI:

ALII:

QR722_09955(fdhF)

CHIZ:

CFON:

SUTT:

SUTK:

CCO:	CCC13826_0200(fdhH)

CCOC:

CCON33237_0536(fdhH)

CGRA:

CGRAC_1361(fdhH)

CPIN:

CPIN18020_0262(fdhH)

CGEO:

CGEO_1649(fdhH)

CMUC:

CMCT_1635(fdhH)

CSHO:

CSHOW_0456(fdhH)

CCAQ:

CCAL_0266(fdhH)

SULS:

Sdiek1_1063

SULJ:

SJPD1_1032

AHS:	AHALO_1025(fdhH)

AMAR:

AMRN_1963(fdhH)

ACAA:

ACAN_2017(fdhH)

CMED:

FE773_08405

GBZ:	JZM60_10225(fdhF)

GEO:

Geob_0149

PCA:

Pcar_2701

PPD:	Ppro_2790 Ppro_2799

DVE:	DESUT3_05340

DDN:	DND132_2347

BWS:	WCP94_000051 WCP94_000257

DOA:	AXF15_00785

DSF:

UWK_03159

DLM:	DPPLL_12850 DPPLL_25480

DAT:	HRM2_11880(fdhA3) HRM2_14370(fdhA4)

DOV:	DSCO28_26720 DSCO28_48980 DSCO28_65560

DWD:	DSCW_21250

DML:	Dmul_02420

DMP:

FAK_06900

FPES:

NXS98_05995

PBAP:

Pla133_02470(fdhF)

BPIT:

BPIT_11130

DTE:	Dester_1341

EFS:

ENE:

CSQ:

DSY:

DHD:

DDH:	Desde_2607

FWA:	DCMF_16640

TJR:	TherJR_2471

DGI:	Desgi_0611 Desgi_0612

DBC:	MFMK1_001222(fdhF)

BACG:

D2962_12430

AAR:	Acear_0635

CPOR:

BED41_00295

MJA:

MJ_0006

MFE:	Mefer_0653 Mefer_1009

MVU:	Metvu_0711 Metvu_1163

MFS:	MFS40622_0207 MFS40622_0236

MJH:	JH146_0247 JH146_0280

MESG:

MLAUSG7_0307 MLAUSG7_0838

MESA:

MLASG1_0357 MLASG1_1372

MIG:	Metig_0762 Metig_1192

MMP:	MMP0138(fdhA) MMP1298(fdhA)

MMQ:	MmarC5_0295 MmarC5_1538

MMX:	MmarC6_0814 MmarC6_1376

MMZ:	MmarC7_0543 MmarC7_1137

MMAK:

MMKA1_01430(fdhA) MMKA1_15230(fdhA)

MMAO:

MMOS7_01690(fdhA) MMOS7_14050(fdhA)

MMAD:

MMJJ_09470(fdhF_2) MMJJ_15440(fdhA2)

MAE:

Maeo_0578

MVN:	Mevan_0608

MVO:	Mvol_0703 Mvol_0819

MOK:	Metok_0188 Metok_1223

MTH:

MTH_1140

MMG:	MTBMA_c15230(fdhA2)

MWO:	MWSIV6_0697(fdhA5) MWSIV6_1532

METC:

MTCT_1033 MTCT_1439

METE:

tca_01096

METK:

FVF72_05885

MTHM:

FZP57_03345 FZP57_07335

METJ:

FZP68_03340

MST:	Msp_0640(fdhA)

METB:

AW729_02760

MEIS:

PXD04_07905

MRU:	mru_0333(fdhA1) mru_2074(fdhA2)

MSI:	Msm_1404 Msm_1463

MEB:	Abm4_0131(fdhA) Abm4_0855(fdhA2)

MMIL:

sm9_1350(flpA) sm9_2183(fdhA)

MEYE:

TL18_01670

MOL:	YLM1_0030 YLM1_1106 YLM1_1540

MARO:

MarbSA_13560

METV:

K4897_04275(fdhF) K4897_08140

MEW:	MSWAN_0116

METH:

MBMB1_0104(fdhA)

MFC:	BRM9_0168(fdhA)

MFI:	DSM1535_1486(fdhA)

MCUB:

MCBB_2028(fdhA)

MSUB:

METN:

METT:

METO:

MFEG:

GCM10025860_09600 GCM10025860_11680 GCM10025860_14950

MRID:

FGU46_06255

MFV:	Mfer_0694 Mfer_0726

MKA:

FPL:

PAB:

PYN:

PYA:	PYCH_11030

PYS:

Py04_1234

PYC:	TQ32_05240 TQ32_09760 TQ32_10390

TKO:	TK0214 TK2076

TON:	TON_0281 TON_0539 TON_1563

TGA:	TGAM_0065(mhy2A) TGAM_0249(fdh1A) TGAM_0631(fdhA)

TSI:

TSIB_0067

TBA:	TERMP_00460

THE:	GQS_02455 GQS_05815

THA:	TAM4_1110 TAM4_610

THM:	CL1_0794 CL1_1148

TLT:	OCC_06696 OCC_12236

THS:	TES1_0096 TES1_1496

TNU:	BD01_0120 BD01_1760 BD01_1899

TEU:	TEU_00760 TEU_06125

TGY:	X802_03675

THV:	ADU37_CDS22830

TCH:	CHITON_0827 CHITON_1559

TPEP:

A0127_02945 A0127_09455

TPIE:

A7C91_04320 A7C91_06945 A7C91_08750

TGG:	A3K92_08040

TCE:	A3L02_01395

TBS:	A3L01_00065

THH:	CDI07_00115

TSL:	A3L11_03805

TTD:	A3L14_01350

TPRF:

A3L09_08935

TRL:	A3L10_09785

TPAF:

A3L08_06500

THY:	A3L12_01285

TIC:	FH039_02070 FH039_05990

TCQ:	TIRI35C_0225(fdhA)

THEM:

FPV09_03360 FPV09_07205

THEI:

K1720_03400(fdhF)

TAGG:

NF865_01295

PPAC:

PAP_03230

MBA:	Mbar_A1561

MBY:	MSBRM_2896

MBW:	MSBRW_3169

MBAR:

MSBR2_2855

MMAC:

MSMAC_2582

METM:

MSMTP_1177

MEF:	MSWH1_2086

MEQ:	MSWHS_3009

MHOR:

MHAB:

MPY:

MSEB:

MEHF:

MEES:

MTP:

MCJ:	MCON_2237(fdhA)

MHI:	Mhar_0325 Mhar_1282 Mhar_1628 Mhar_1941

MHU:	Mhun_1813 Mhun_1833 Mhun_2021 Mhun_2023 Mhun_3238

MRTJ:

KHC33_00530(fdhF) KHC33_05240(fdhF) KHC33_05340(fdhF) KHC33_10640(fdhF) KHC33_14310(fdhF) KHC33_14320(fdhF)

MLA:	Mlab_0463 Mlab_1057

MEM:	Memar_1045 Memar_1379 Memar_1381 Memar_1624

MBG:	BN140_0842(fdhA) BN140_1327(fdhA1) BN140_1329(fdhA2) BN140_2198(fdhA3)

MEMA:

MMAB1_1106(fdhA) MMAB1_1689(fdhA) MMAB1_2914(fdhA)

MCHK:

MchiMG62_11700 MchiMG62_15760 MchiMG62_15780 MchiMG62_17900

MSUM:

OH143_01120(fdhF) OH143_01130(fdhF) OH143_02330(fdhF) OH143_11950(fdhF)

MRC:	R6Y96_05800(fdhF) R6Y96_06465(fdhF)

MPI:	Mpet_1036 Mpet_1048 Mpet_1628

MEND:

L6E24_11840(fdhF)

MEFW:

F1737_05530 F1737_05555

MAQE:

RJ40_03450

MFK:	E2N92_03505

MOU:	OU421_01580(fdhF)

MESB:

L1S32_05095(fdhF) L1S32_07140

MANQ:

L1994_04160 L1994_05190(fdhF)

MBN:

Mboo_0923

MFO:	Metfor_0647 Metfor_0650 Metfor_0677

MPL:	Mpal_0196 Mpal_0578

MPD:	MCP_1570(fdhA-1) MCP_2407(fdhA-2)

MEZ:	Mtc_2125(fdhA)

RCI:	RCIX1628(fdhA-1) RCIX2373(fdhA-2) RCIX2378(fdhA-3) RRC251(fdhA-4)

MJK:	N2V74_00365 N2V74_05490

HHB:	Hhub_2705(fdhA)

HNO:	LT974_10300(fdhF)

HANX:

ABSL23_13375(fdhF)

HALH:

HTSR_1736(fdhA) HTSR_1740(fdhA2)

HHSR:

HSR6_1802(fdoG2) HSR6_1806(fdoG3)

HJE:	HacjB3_08400

HARA:

AArcS_1771(yjgC)

HHI:	HAH_5041(fdhA)

HHN:	HISP_18370

HAB:	SG26_17860

HALI:

BV210_11270

SSAI:

N0B31_21065(fdhF)

NMO:	Nmlp_2714(fdhA)

HAQ:	DU484_02900 DU484_17375

HAJ:	DU500_00415 DU500_03330

HAER:

DU502_01085 DU502_15800

HRA:	EI982_15430

HLM:	DV707_15380

HLA:

Hlac_1238

HALP:

DOS48_05995

HALB:

EKH57_00640 EKH57_15440

HEZZ:

EO776_02790

HALQ:

Hrr1229_013155(fdhF)

HSS:	J7656_05930(fdhF) J7656_13980(fdhF)

HSAI:

HPS36_11145(fdhF)

HAZZ:

KI388_08835(fdhF)

SRUB:

C2R22_06085

HALU:

HUG12_20610(fdhF)

HRM:	K6T25_13375(fdhF)

HDF:	AArcSl_0465(fdoG) AArcSl_1489(fdoG2)

HAAA:

AArcCO_2264(yjgC)

HTU:

Htur_0685

HSAL:

JMJ58_05665(fdhF)

HXA:	Halxa_0835

NPE:	Natpe_2904

NAY:	HYG81_16490(fdhF)

HALY:

HYG82_00570(fdhF)

HJT:	DVR14_04645

NGE:	Natgr_0651 Natgr_2918

HLC:	CHINAEXTREME08970

NOU:	Natoc_1422

NAJ:	B1756_07310 B1756_16645

NAG:	AArcMg_0491

NAN:	AArc1_0503

NARA:

QQ977_13275(fdhF)

NBG:	DV706_17960

NAS:	GCU68_09540

TAC:

Ta0045

TVO:	TVG0003442(TVG0003442)

FAC:	FACI_IFERC01G0549

FAI:

FAD_1471

ABI:

Aboo_0808

SHC:	Shell_0724

IHO:

Igni_1101

IIS:

EYM_07315

PFM:	Pyrfu_0745

AHO:

AAMB:

VDI:

TPE:

FFO:	FFONT_1292

» show all

Reference

PMID:18791018

Authors

Lupa B, Hendrickson EL, Leigh JA, Whitman WB

Title

Formate-dependent H2 production by the mesophilic methanogen Methanococcus maripaludis.

Journal

Appl Environ Microbiol 74:6584-90 (2008)
DOI:10.1128/AEM.01455-08

Reference

PMID:24039260

Authors

Costa KC, Lie TJ, Xia Q, Leigh JA

Title

VhuD facilitates electron flow from H2 or formate to heterodisulfide reductase in Methanococcus maripaludis.

Journal

J Bacteriol 195:5160-5 (2013)
DOI:10.1128/JB.00895-13

LinkDB

ORTHOLOGY: K00125

Entry

K00125                      KO

Symbol

fdhB

Name

formate dehydrogenase (coenzyme F420) beta subunit [EC:1.17.98.3 1.8.98.6]

Pathway

map00680

Methane metabolism

map01100

Metabolic pathways

map01120

Microbial metabolism in diverse environments

map01200

Carbon metabolism

Module

M00356

Methanogenesis, methanol => methane

M00357

Methanogenesis, acetate => methane

M00563

Methanogenesis, methylamine/dimethylamine/trimethylamine => methane

M00567

Methanogenesis, CO2 => methane

Reaction

R10797

formate:coenzyme-F420 oxidoreductase

R11944

coenzyme B,coenzyme M,ferredoxin:formate oxidoreductase

Brite

KEGG Orthology (KO) [BR:ko00001]
09100 Metabolism
  09102 Energy metabolism
   00680 Methane metabolism
    K00125  fdhB; formate dehydrogenase (coenzyme F420) beta subunit
Enzymes [BR:ko01000]
1. Oxidoreductases
  1.8  Acting on a sulfur group of donors
   1.8.98  With other, known, physiological acceptors
    1.8.98.6  formate:CoB-CoM heterodisulfide,ferredoxin reductase
     K00125  fdhB; formate dehydrogenase (coenzyme F420) beta subunit
  1.17  Acting on CH or CH2 groups
   1.17.98  With other, known, physiological acceptors
    1.17.98.3  formate dehydrogenase (coenzyme F420)
     K00125  fdhB; formate dehydrogenase (coenzyme F420) beta subunit

Other DBs

COG:	COG1035 COG1139

GO:

0043794

Genes

TMB:	Thimo_0255

NPV:	OHM77_12825

SLAC:

SKTS_06260

BLAG:

BLTE_14910

RAP:

RHOA_4366

GSU:	GSU0088(hdrD)

GSK:	KN400_0063(hdrD)

GME:	Gmet_3428(hdrD)

GPI:	GPICK_06770 GPICK_15705

GAO:	A2G06_16000

GBZ:	JZM60_06200

GEM:

GM21_0027

GEB:	GM18_0024 GM18_3241

GEF:	FO488_02625

GEV:	GSVR_37980(hdrD)

GEC:	GEO60473_29700

GUR:

Gura_0147

GEO:	Geob_3720(hdrD)

TAMM:

GEAMG1_2371(hdrD)

GBM:	Gbem_0028(hdrD)

GBN:	GEOBRER4_00260(hdrD)

GER:	KP004_19730

GSUB:

KP001_16585

GNT:	KP003_19720

GPL:	M1B72_00020

DFL:	DFE_0758 DFE_2489

DMA:

DMR_01940

DCB:	C3Y92_00575

DGG:	DGI_1052(floxC)

DDE:	Dde_1211 Dde_3528

DMS:	E8L03_02850

DSA:	Desal_0598

DHY:	DESAM_21328

DAF:	Desaf_1864 Desaf_2879

DAS:

Daes_0781

DPI:

BN4_11862

DEJ:	AWY79_07305

PSEL:

GM415_15615

DDN:	DND132_1390

PPOR:

PNW:

PTHE:

PMEN:

DSX:	GD604_05900 GD604_12445

DSD:	GD606_00175 GD606_07840 GD606_11690

DBA:

Dbac_2346

DOA:	AXF15_11190

DESU:

NLA06_01255

DRT:

Dret_0419

DLM:	DPPLL_25470(hdrD)

ECOM:

QTN59_02640

EAJ:	Q3M24_19845

ELAX:

KKHLCK_04500

DAL:

Dalk_5062

DAT:	HRM2_22080(fdhB5) HRM2_22740(fdhB6)

DTO:	TOL2_C17890 TOL2_C24200

DOV:	DSCO28_27580 DSCO28_27830 DSCO28_36090

DWD:	DSCW_47550 DSCW_60930 DSCW_61120

DALK:

DSCA_54490 DSCA_54710 DSCA_62380(hdrD)

DVR:	Dvar_21620

DMM:	dnm_011400 dnm_084970

DEK:	DSLASN_12960

DTI:	Desti_4266

SFU:	Sfum_0822 Sfum_1972

DAX:	FDQ92_05250 FDQ92_10770

DBR:

Deba_2134

TID:	Thein_2071

TAV:	G4V39_00755 G4V39_00795

DAW:	HS1_002392 HS1_002396

TBE:	Trebr_2032

TTK:	TST_1517 TST_1524

CEU:	A7L45_12260

CTHD:

RCH:

RUM:

SMAN:

HMB:	ACI0CA_03150

SWO:

Swol_0400

SALQ:

SYNTR_0266

SGY:

Sgly_1898

DOR:	Desor_2815

DAI:	Desaci_2628

DMI:	Desmer_2224

DCA:	Desca_0132 Desca_0138 Desca_0819

PTH:	PTH_1407(RnfC)

DRM:	Dred_0139 Dred_0146 Dred_1330

DRU:	Desru_0208 Desru_0215 Desru_2695

DFG:	B0537_01500 B0537_01535 B0537_06305 B0537_08745

DAE:	Dtox_1279 Dtox_1355

DKU:	Desku_0190 Desku_1432 Desku_1482

TFR:	BR63_02785 BR63_09665

FWA:	DCMF_10300 DCMF_24650

TJR:	TherJR_0939 TherJR_1951

DGI:	Desgi_2044

HMO:

HM1_2010

HCV:	FTV88_1706

CTHM:

CFE_1626

BYL:	A4V09_16705

BPRO:

PMF13cell1_01674

BCOC:

BLCOC_07560

BLAB:

EYS05_14060

BWX:	NQ550_09275

BMAS:

BLAR:

BLUI:

MFOR:

AHAD:

CMIU:

DAU:

CHY:

ADG:

MAS:	Mahau_0897

NTH:	Nther_2660

ABAT:

CFX1CAM_1232

PSUB:

Pelsub_P0987

FVA:	FV113G1_17800

FUL:	C4N20_13335

IPO:	Ilyop_2562

MJA:

MJ_0005

MFE:	Mefer_0654

MVU:	Metvu_0710

MFS:	MFS40622_0206 MFS40622_0235

MJH:	JH146_0281

MESG:

MLAUSG7_0837(fdhB)

MESA:

MLASG1_0358(fdhB)

MIG:	Metig_1191

MMP:	MMP0139(fdhB) MMP1297(fdhB)

MMQ:	MmarC5_0296 MmarC5_1537

MMX:	MmarC6_0813 MmarC6_1377

MMZ:	MmarC7_0542 MmarC7_1138

MMD:	GYY_00710 GYY_07320

MMAK:

MMKA1_01440(fdhB) MMKA1_15220(fdhB)

MMAO:

MMOS7_01700(fdhB) MMOS7_14040(fdhB)

MMAD:

MMJJ_09460 MMJJ_15460(fdhB1)

MAE:

Maeo_0577

MVN:	Mevan_0607 Mevan_1144

MVO:	Mvol_0702 Mvol_0818

MOK:	Metok_0189 Metok_1224

MTH:

MTH_1139

MMG:	MTBMA_c15220(fdhB)

MWO:	MWSIV6_0698(fdhB1) MWSIV6_1531(fdhB3)

MTEE:

METC:

METE:

METK:

MTHM:

FZP57_03350 FZP57_07330

METJ:

FZP68_03345

MST:	Msp_0639(fdhB)

METB:

AW729_02755

MEIS:

PXD04_07910

MRU:	mru_0334(fdhB1) mru_2075(fdhB2)

MSI:	Msm_1405 Msm_1462

MEB:	Abm4_0132(fdhB) Abm4_0854(fdhB2)

MMIL:

sm9_1349(flpB) sm9_2182(fdhB)

MEYE:

TL18_00925 TL18_01675

MOL:	YLM1_0031 YLM1_1105 YLM1_1539

MARO:

MarbSA_13570

METV:

K4897_04270 K4897_08135

MEW:	MSWAN_0113 MSWAN_0115

METH:

MBMB1_0103(fdhB)

MFC:	BRM9_0167(fdhB)

MFI:	DSM1535_1485(fdhB)

MCUB:

MCBB_2029(fdhB1) MCBB_2031(fdhB3)

MSUB:

BK009_10240 BK009_10250

METN:

BK008_05865 BK008_05875

METT:

CIT01_01080

METO:

CIT02_06505

MFEG:

GCM10025860_09590 GCM10025860_11690 GCM10025860_14940

MRID:

FGU46_06240 FGU46_06250

MFV:	Mfer_0695 Mfer_0725

MKA:

MK0322

AVE:	Arcve_2107

AST:	Asulf_01818

MBA:	Mbar_A1562

MBY:	MSBRM_2895

MBW:	MSBRW_3168

MBAR:

MMAC:

MHOR:

MHAB:

MPY:

MSEB:

MEHF:

MEES:

MHI:

MHU:	Mhun_1814 Mhun_1832 Mhun_2020 Mhun_2022 Mhun_3237

MRTJ:

KHC33_00535 KHC33_05245 KHC33_05335 KHC33_10635 KHC33_14315 KHC33_14325

MLA:	Mlab_0462 Mlab_1058

MEM:	Memar_1044 Memar_1378 Memar_1380 Memar_1623

MBG:	BN140_0841(fdhB) BN140_1328(fdhB1) BN140_1330(fdhB2) BN140_2197(fdhB3)

MEMA:

MMAB1_1105(fdhB) MMAB1_1690(fdhB) MMAB1_2913(fdhB)

MCHK:

MchiMG62_11690 MchiMG62_15750 MchiMG62_15770 MchiMG62_17910

MSUM:

OH143_01115 OH143_01125 OH143_02325 OH143_11945

MRC:	R6Y96_05805 R6Y96_06470

MPI:	Mpet_1035 Mpet_1047 Mpet_1627

MEND:

L6E24_11845

MEFW:

F1737_05525 F1737_05550

MAQE:

RJ40_03445

MFK:	E2N92_03500

MOU:	OU421_01585 OU421_03545

MESB:

L1S32_05100 L1S32_07145

MANQ:

L1994_04165 L1994_05185

MBN:

Mboo_0924

MFO:	Metfor_0648 Metfor_0651 Metfor_0678

MESK:

METHP_01621

MPL:	Mpal_0195 Mpal_0577

MPD:	MCP_1569(fdhB-1) MCP_2406(fdhB-2)

MEZ:	Mtc_2124(fdhB)

RCI:	RCIX1630(fdhB-1) RCIX2372(fdhB-2) RCIX2376(fdhB-3) RRC252(fdhB-4)

MJK:	N2V74_00370 N2V74_01825 N2V74_04035 N2V74_05485

ABI:

Aboo_0980

ACF:	AciM339_1138

LOKI:

Lokiarch_00320(fdhB)

LOB:	NEF87_000179

PSYT:

DSAG12_02784 DSAG12_03719

» show all

Reference

PMID:18791018

Authors

Lupa B, Hendrickson EL, Leigh JA, Whitman WB

Title

Formate-dependent H2 production by the mesophilic methanogen Methanococcus maripaludis.

Journal

Appl Environ Microbiol 74:6584-90 (2008)
DOI:10.1128/AEM.01455-08

Reference

PMID:24039260

Authors

Costa KC, Lie TJ, Xia Q, Leigh JA

Title

VhuD facilitates electron flow from H2 or formate to heterodisulfide reductase in Methanococcus maripaludis.

Journal

J Bacteriol 195:5160-5 (2013)
DOI:10.1128/JB.00895-13

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