KEGG   PATHWAY: nau00270
Entry
nau00270                    Pathway                                
Name
Cysteine and methionine metabolism - Nicotiana attenuata (coyote tobacco)
Description
Cysteine and methionine are sulfur-containing amino acids. Cysteine is synthesized from serine through different pathways in different organism groups. In bacteria and plants, cysteine is converted from serine (via acetylserine) by transfer of hydrogen sulfide [MD:M00021]. In animals, methionine-derived homocysteine is used as sulfur source and its condensation product with serine (cystathionine) is converted to cysteine [MD:M00338]. Cysteine is metabolized to pyruvate in multiple routes. Methionine is an essential amino acid, which animals cannot synthesize. In bacteria and plants, methionine is synthesized from aspartate [MD:M00017]. S-Adenosylmethionine (SAM), synthesized from methionine and ATP, is a methyl group donor in many important transfer reactions including DNA methylation for regulation of gene expression. SAM may also be used to regenerate methionine in the methionine salvage pathway [MD:M00034].
Class
Metabolism; Amino acid metabolism
Pathway map
nau00270  Cysteine and methionine metabolism
nau00270

Module
nau_M00021  Cysteine biosynthesis, serine => cysteine [PATH:nau00270]
nau_M00034  Methionine salvage pathway [PATH:nau00270]
nau_M00368  Ethylene biosynthesis, methionine => ethylene [PATH:nau00270]
Other DBs
GO: 0006534 0006555
Organism
Nicotiana attenuata (coyote tobacco) [GN:nau]
Gene
109213540  serine acetyltransferase 2-like [KO:K00640] [EC:2.3.1.30]
109211171  serine acetyltransferase 1, chloroplastic-like [KO:K00640] [EC:2.3.1.30]
109225207  serine acetyltransferase 5 [KO:K00640] [EC:2.3.1.30]
109225518  serine acetyltransferase 1, chloroplastic-like [KO:K00640] [EC:2.3.1.30]
109244706  cysteine synthase [KO:K01738] [EC:2.5.1.47]
109237848  cysteine synthase-like isoform X1 [KO:K01738] [EC:2.5.1.47]
109236512  cysteine synthase, chloroplastic/chromoplastic [KO:K01738] [EC:2.5.1.47]
109229022  cysteine synthase 2 [KO:K01738] [EC:2.5.1.47]
109231094  bifunctional L-3-cyanoalanine synthase/cysteine synthase 1, mitochondrial [KO:K13034] [EC:2.5.1.47 4.4.1.9]
109243119  bifunctional L-3-cyanoalanine synthase/cysteine synthase 2, mitochondrial [KO:K13034] [EC:2.5.1.47 4.4.1.9]
109218926  cystathionine beta-lyase, chloroplastic-like [KO:K01760] [EC:4.4.1.13]
109239251  cystathionine beta-lyase, chloroplastic [KO:K01760] [EC:4.4.1.13]
109227184  homocysteine S-methyltransferase 2-like [KO:K00547] [EC:2.1.1.10]
109218873  homocysteine S-methyltransferase 2-like [KO:K00547] [EC:2.1.1.10]
109221165  homocysteine S-methyltransferase 1 [KO:K00547] [EC:2.1.1.10]
109213532  5-methyltetrahydropteroyltriglutamate-- homocysteine methyltransferase-like [KO:K00549] [EC:2.1.1.14]
109238895  5-methyltetrahydropteroyltriglutamate-- homocysteine methyltransferase-like [KO:K00549] [EC:2.1.1.14]
109220679  5-methyltetrahydropteroyltriglutamate-- homocysteine methyltransferase-like [KO:K00549] [EC:2.1.1.14]
109238602  5-methyltetrahydropteroyltriglutamate-- homocysteine methyltransferase [KO:K00549] [EC:2.1.1.14]
109233364  S-adenosylmethionine synthase 1 [KO:K00789] [EC:2.5.1.6]
109210093  S-adenosylmethionine synthase 2-like [KO:K00789] [EC:2.5.1.6]
109210603  S-adenosylmethionine synthase 3-like [KO:K00789] [EC:2.5.1.6]
109229653  S-adenosylmethionine synthase 3 [KO:K00789] [EC:2.5.1.6]
109235709  S-adenosylmethionine synthase 2 [KO:K00789] [EC:2.5.1.6]
109214247  S-adenosylmethionine synthase 2 [KO:K00789] [EC:2.5.1.6]
109243880  S-adenosylmethionine decarboxylase proenzyme 4-like [KO:K01611] [EC:4.1.1.50]
109232750  S-adenosylmethionine decarboxylase proenzyme 4-like [KO:K01611] [EC:4.1.1.50]
109231064  S-adenosylmethionine decarboxylase proenzyme-like [KO:K01611] [EC:4.1.1.50]
109212838  S-adenosylmethionine decarboxylase proenzyme [KO:K01611] [EC:4.1.1.50]
109210188  S-adenosylmethionine decarboxylase proenzyme-like [KO:K01611] [EC:4.1.1.50]
109217294  S-adenosylmethionine decarboxylase proenzyme-like [KO:K01611] [EC:4.1.1.50]
109217298  S-adenosylmethionine decarboxylase proenzyme-like [KO:K01611] [EC:4.1.1.50]
109234538  S-adenosylmethionine decarboxylase proenzyme-like [KO:K01611] [EC:4.1.1.50]
109213904  spermidine synthase 1 [KO:K00797] [EC:2.5.1.16]
109235629  spermidine synthase [KO:K00797] [EC:2.5.1.16]
109223522  spermine synthase [KO:K00797] [EC:2.5.1.16]
109233844  nicotianamine synthase-like [KO:K05953] [EC:2.5.1.43]
109223070  nicotianamine synthase [KO:K05953] [EC:2.5.1.43]
109217211  5'-methylthioadenosine/S-adenosylhomocysteine nucleosidase 2-like [KO:K01244] [EC:3.2.2.16]
109216992  methylthioribose kinase [KO:K00899] [EC:2.7.1.100]
109209621  methylthioribose-1-phosphate isomerase isoform X1 [KO:K08963] [EC:5.3.1.23]
109230742  probable bifunctional methylthioribulose-1-phosphate dehydratase/enolase-phosphatase E1 1 [KO:K16054] [EC:4.2.1.109 3.1.3.77]
109229163  probable bifunctional methylthioribulose-1-phosphate dehydratase/enolase-phosphatase E1 1 isoform X1 [KO:K16054] [EC:4.2.1.109 3.1.3.77]
109221219  1,2-dihydroxy-3-keto-5-methylthiopentene dioxygenase 2 [KO:K08967] [EC:1.13.11.53 1.13.11.54]
109229678  1,2-dihydroxy-3-keto-5-methylthiopentene dioxygenase 2 [KO:K08967] [EC:1.13.11.53 1.13.11.54]
109212046  1,2-dihydroxy-3-keto-5-methylthiopentene dioxygenase 1 [KO:K08967] [EC:1.13.11.53 1.13.11.54]
109237084  uncharacterized protein LOC109237084 [KO:K23977] [EC:2.6.1.117]
109239085  tyrosine aminotransferase-like [KO:K00815] [EC:2.6.1.5]
109230869  probable aminotransferase TAT2 [KO:K00815] [EC:2.6.1.5]
109227275  tyrosine aminotransferase-like [KO:K00815] [EC:2.6.1.5]
109227276  tyrosine aminotransferase-like [KO:K00815] [EC:2.6.1.5]
109223541  probable aminotransferase TAT2 [KO:K00815] [EC:2.6.1.5]
109240631  probable aminotransferase TAT2 [KO:K00815] [EC:2.6.1.5]
109221888  probable aminotransferase TAT2 [KO:K00815] [EC:2.6.1.5]
109231153  uncharacterized protein LOC109231153 [KO:K00837] [EC:2.6.1.-]
109244641  methionine gamma-lyase-like [KO:K01761] [EC:4.4.1.11]
109219429  methionine gamma-lyase-like [KO:K01761] [EC:4.4.1.11]
109241305  DNA (cytosine-5)-methyltransferase CMT2 [KO:K00558] [EC:2.1.1.37]
109205822  DNA (cytosine-5)-methyltransferase CMT3-like [KO:K00558] [EC:2.1.1.37]
109235780  DNA (cytosine-5)-methyltransferase CMT3-like [KO:K00558] [EC:2.1.1.37]
109240115  DNA (cytosine-5)-methyltransferase 1B-like [KO:K00558] [EC:2.1.1.37]
109217400  uncharacterized protein LOC109217400 [KO:K17398] [EC:2.1.1.37]
109219843  adenosylhomocysteinase-like [KO:K01251] [EC:3.13.2.1]
109243151  adenosylhomocysteinase [KO:K01251] [EC:3.13.2.1]
109220219  1-aminocyclopropane-1-carboxylate synthase-like [KO:K20772] [EC:4.4.1.14]
109237500  1-aminocyclopropane-1-carboxylate synthase-like [KO:K20772] [EC:4.4.1.14]
109241083  1-aminocyclopropane-1-carboxylate synthase-like [KO:K20772] [EC:4.4.1.14]
109241084  1-aminocyclopropane-1-carboxylate synthase-like [KO:K20772] [EC:4.4.1.14]
109205644  1-aminocyclopropane-1-carboxylate synthase [KO:K20772] [EC:4.4.1.14]
109229811  1-aminocyclopropane-1-carboxylate synthase 3-like [KO:K01762] [EC:4.4.1.14]
109239471  1-aminocyclopropane-1-carboxylate synthase 3-like [KO:K01762] [EC:4.4.1.14]
109210465  1-aminocyclopropane-1-carboxylate synthase 3-like [KO:K01762] [EC:4.4.1.14]
109220530  1-aminocyclopropane-1-carboxylate synthase 7-like [KO:K01762] [EC:4.4.1.14]
109223202  1-aminocyclopropane-1-carboxylate synthase 7-like [KO:K01762] [EC:4.4.1.14]
109218840  1-aminocyclopropane-1-carboxylate synthase 3 [KO:K01762] [EC:4.4.1.14]
109205942  1-aminocyclopropane-1-carboxylate oxidase-like [KO:K05933] [EC:1.14.17.4]
109218050  1-aminocyclopropane-1-carboxylate oxidase 5-like [KO:K05933] [EC:1.14.17.4]
109228672  1-aminocyclopropane-1-carboxylate oxidase [KO:K05933] [EC:1.14.17.4]
109228673  1-aminocyclopropane-1-carboxylate oxidase-like [KO:K05933] [EC:1.14.17.4]
109240562  1-aminocyclopropane-1-carboxylate oxidase 5 [KO:K05933] [EC:1.14.17.4]
109220508  1-aminocyclopropane-1-carboxylate oxidase 1 [KO:K05933] [EC:1.14.17.4]
109223193  1-aminocyclopropane-1-carboxylate oxidase-like [KO:K05933] [EC:1.14.17.4]
109229011  1-aminocyclopropane-1-carboxylate oxidase [KO:K05933] [EC:1.14.17.4]
109222014  1-aminocyclopropane-1-carboxylate oxidase-like [KO:K05933] [EC:1.14.17.4]
109226594  1-aminocyclopropane-1-carboxylate oxidase 1 [KO:K05933] [EC:1.14.17.4]
109222454  aspartokinase 1, chloroplastic-like [KO:K00928] [EC:2.7.2.4]
109243291  aspartokinase 2, chloroplastic-like isoform X1 [KO:K00928] [EC:2.7.2.4]
109216117  bifunctional aspartokinase/homoserine dehydrogenase 1, chloroplastic-like [KO:K12524] [EC:2.7.2.4 1.1.1.3]
109243638  bifunctional aspartokinase/homoserine dehydrogenase, chloroplastic-like [KO:K12524] [EC:2.7.2.4 1.1.1.3]
109243293  uncharacterized protein LOC109243293 [KO:K00133] [EC:1.2.1.11]
109222320  uncharacterized protein LOC109222320 [KO:K00133] [EC:1.2.1.11]
109230850  cystathionine gamma-synthase 1, chloroplastic-like [KO:K01739] [EC:2.5.1.48]
109211558  cystathionine gamma-synthase 1, chloroplastic-like [KO:K01739] [EC:2.5.1.48]
109213315  cystathionine gamma-synthase 1, chloroplastic-like [KO:K01739] [EC:2.5.1.48]
109244694  branched-chain-amino-acid aminotransferase 3, chloroplastic-like [KO:K00826] [EC:2.6.1.42]
109211373  branched-chain-amino-acid aminotransferase 2, chloroplastic-like [KO:K00826] [EC:2.6.1.42]
109233330  branched-chain-amino-acid aminotransferase 2, chloroplastic-like isoform X1 [KO:K00826] [EC:2.6.1.42]
109239465  branched-chain-amino-acid aminotransferase 2, chloroplastic-like isoform X1 [KO:K00826] [EC:2.6.1.42]
109228853  branched-chain-amino-acid aminotransferase 3, chloroplastic-like [KO:K00826] [EC:2.6.1.42]
109219791  branched-chain-amino-acid aminotransferase 2, chloroplastic-like [KO:K00826] [EC:2.6.1.42]
109207780  branched-chain-amino-acid aminotransferase 2, chloroplastic-like [KO:K00826] [EC:2.6.1.42]
109210516  alanine--glyoxylate aminotransferase 2 homolog 3, mitochondrial-like [KO:K00827] [EC:2.6.1.44 2.6.1.40]
109209547  alanine--glyoxylate aminotransferase 2 homolog 1, mitochondrial isoform X1 [KO:K00827] [EC:2.6.1.44 2.6.1.40]
109233404  alanine--glyoxylate aminotransferase 2 homolog 3, mitochondrial-like [KO:K00827] [EC:2.6.1.44 2.6.1.40]
109242261  glutamate--cysteine ligase, chloroplastic [KO:K01919] [EC:6.3.2.2]
109221207  glutathione synthetase, chloroplastic isoform X1 [KO:K21456] [EC:6.3.2.3]
109234188  aspartate aminotransferase, cytoplasmic [KO:K14454] [EC:2.6.1.1]
109241053  aspartate aminotransferase, cytoplasmic [KO:K14454] [EC:2.6.1.1]
109225049  aspartate aminotransferase, mitochondrial [KO:K14455] [EC:2.6.1.1]
109233753  aspartate aminotransferase, mitochondrial-like [KO:K14455] [EC:2.6.1.1]
109214188  aspartate aminotransferase, chloroplastic [KO:K00811] [EC:2.6.1.1]
109234106  thiosulfate/3-mercaptopyruvate sulfurtransferase 1, mitochondrial-like [KO:K01011] [EC:2.8.1.1 2.8.1.2]
109238458  thiosulfate/3-mercaptopyruvate sulfurtransferase 1, mitochondrial-like [KO:K01011] [EC:2.8.1.1 2.8.1.2]
109232867  L-lactate dehydrogenase B-like [KO:K00016] [EC:1.1.1.27]
109216926  L-lactate dehydrogenase B-like [KO:K00016] [EC:1.1.1.27]
109228643  D-cysteine desulfhydrase 2, mitochondrial isoform X1 [KO:K05396] [EC:4.4.1.15]
109208697  bifunctional D-cysteine desulfhydrase/1-aminocyclopropane-1-carboxylate deaminase, mitochondrial isoform X1 [KO:K05396] [EC:4.4.1.15]
109224794  L-cysteine desulfhydrase [KO:K22207] [EC:4.4.1.28]
109243283  probable L-cysteine desulfhydrase, chloroplastic [KO:K22207] [EC:4.4.1.28]
109237246  malate dehydrogenase, cytoplasmic-like isoform X1 [KO:K00025] [EC:1.1.1.37]
109239064  malate dehydrogenase-like [KO:K00025] [EC:1.1.1.37]
109220558  malate dehydrogenase [KO:K00025] [EC:1.1.1.37]
109221966  malate dehydrogenase-like [KO:K00025] [EC:1.1.1.37]
109213586  malate dehydrogenase, glyoxysomal [KO:K00026] [EC:1.1.1.37]
109211238  malate dehydrogenase, chloroplastic isoform X1 [KO:K00026] [EC:1.1.1.37]
109230769  malate dehydrogenase, mitochondrial-like [KO:K00026] [EC:1.1.1.37]
109211734  malate dehydrogenase, glyoxysomal-like [KO:K00026] [EC:1.1.1.37]
109226664  malate dehydrogenase 2, mitochondrial [KO:K00026] [EC:1.1.1.37]
109232416  malate dehydrogenase, chloroplastic-like [KO:K00026] [EC:1.1.1.37]
109221201  D-3-phosphoglycerate dehydrogenase 2, chloroplastic-like [KO:K00058] [EC:1.1.1.95 1.1.1.399]
109214787  D-3-phosphoglycerate dehydrogenase 2, chloroplastic-like [KO:K00058] [EC:1.1.1.95 1.1.1.399]
109224001  D-3-phosphoglycerate dehydrogenase 1, chloroplastic-like [KO:K00058] [EC:1.1.1.95 1.1.1.399]
109210691  D-3-phosphoglycerate dehydrogenase 1, chloroplastic-like [KO:K00058] [EC:1.1.1.95 1.1.1.399]
109233716  D-3-phosphoglycerate dehydrogenase 1, chloroplastic-like [KO:K00058] [EC:1.1.1.95 1.1.1.399]
109213539  phosphoserine aminotransferase 2, chloroplastic-like [KO:K00831] [EC:2.6.1.52]
Compound
C00019  S-Adenosyl-L-methionine
C00021  S-Adenosyl-L-homocysteine
C00022  Pyruvate
C00041  L-Alanine
C00049  L-Aspartate
C00051  Glutathione
C00059  Sulfate
C00065  L-Serine
C00073  L-Methionine
C00094  Sulfite
C00097  L-Cysteine
C00109  2-Oxobutanoate
C00155  L-Homocysteine
C00170  5'-Methylthioadenosine
C00197  3-Phospho-D-glycerate
C00263  L-Homoserine
C00283  Hydrogen sulfide
C00409  Methanethiol
C00441  L-Aspartate 4-semialdehyde
C00491  L-Cystine
C00506  L-Cysteate
C00606  3-Sulfino-L-alanine
C00793  D-Cysteine
C00957  Mercaptopyruvate
C00979  O-Acetyl-L-serine
C01005  O-Phospho-L-serine
C01077  O-Acetyl-L-homoserine
C01118  O-Succinyl-L-homoserine
C01137  S-Adenosylmethioninamine
C01180  4-Methylthio-2-oxobutanoic acid
C01234  1-Aminocyclopropane-1-carboxylate
C01817  L-Homocystine
C01962  Thiocysteine
C02218  Dehydroalanine
C02291  L-Cystathionine
C02356  (S)-2-Aminobutanoate
C02989  L-Methionine S-oxide
C03082  4-Phospho-L-aspartate
C03089  5-Methylthio-D-ribose
C03145  N-Formylmethionine
C03232  3-Phosphonooxypyruvate
C03431  S-Inosyl-L-homocysteine
C03539  S-Ribosyl-L-homocysteine
C04188  S-Methyl-5-thio-D-ribose 1-phosphate
C04582  S-Methyl-5-thio-D-ribulose 1-phosphate
C05324  Nicotianamine
C05524  Aminoacyl-L-methionine
C05526  S-Glutathionyl-L-cysteine
C05527  3-Sulfinylpyruvate
C05528  3-Sulfopyruvate
C05823  3-Mercaptolactate
C05824  S-Sulfo-L-cysteine
C06547  Ethylene
C08276  3-(Methylthio)propanoate
C09306  Sulfur dioxide
C11437  1-Deoxy-D-xylulose 5-phosphate
C11481  HSO3-
C11499  (S)-3-Sulfolactate
C11537  (2R)-3-Sulfolactate
C15606  1,2-Dihydroxy-5-(methylthio)pent-1-en-3-one
C15650  2,3-Diketo-5-methylthiopentyl-1-phosphate
C15651  2-Hydroxy-3-keto-5-methylthiopentenyl-1-phosphate
C18049  N-Acyl-L-homoserine lactone
C19787  5'-S-Methyl-5'-thioinosine
C21015  gamma-L-Glutamyl-L-2-aminobutyrate
C21016  Ophthalmate
C22359  S-Methyl-1-thio-D-xylulose 5-phosphate
Reference
PMID:15102328
  Authors
Sekowska A, Denervaud V, Ashida H, Michoud K, Haas D, Yokota A, Danchin A
  Title
Bacterial variations on the methionine salvage pathway.
  Journal
BMC Microbiol 4:9 (2004)
DOI:10.1186/1471-2180-4-9
Reference
PMID:14551435
  Authors
Ashida H, Saito Y, Kojima C, Kobayashi K, Ogasawara N, Yokota A.
  Title
A functional link between RuBisCO-like protein of Bacillus and photosynthetic RuBisCO.
  Journal
Science 302:286-90 (2003)
DOI:10.1126/science.1086997
Reference
PMID:12022921
  Authors
Sekowska A, Danchin A.
  Title
The methionine salvage pathway in Bacillus subtilis.
  Journal
BMC Microbiol 2:8 (2002)
DOI:10.1186/1471-2180-2-8
Reference
PMID:12670965
  Authors
Berger BJ, English S, Chan G, Knodel MH.
  Title
Methionine regeneration and aminotransferases in Bacillus subtilis, Bacillus cereus, and Bacillus anthracis.
  Journal
J Bacteriol 185:2418-31 (2003)
DOI:10.1128/JB.185.8.2418-2431.2003
Reference
PMID:17169919
  Authors
Goyer A, Collakova E, Shachar-Hill Y, Hanson AD
  Title
Functional characterization of a methionine gamma-lyase in Arabidopsis and its implication in an alternative to the reverse trans-sulfuration pathway.
  Journal
Plant Cell Physiol 48:232-42 (2007)
DOI:10.1093/pcp/pcl055
Reference
PMID:17030798
  Authors
Rebeille F, Jabrin S, Bligny R, Loizeau K, Gambonnet B, Van Wilder V, Douce R, Ravanel S
  Title
Methionine catabolism in Arabidopsis cells is initiated by a gamma-cleavage process and leads to S-methylcysteine and isoleucine syntheses.
  Journal
Proc Natl Acad Sci U S A 103:15687-92 (2006)
DOI:10.1073/pnas.0606195103
Reference
PMID:18625006
  Authors
Pirkov I, Norbeck J, Gustafsson L, Albers E
  Title
A complete inventory of all enzymes in the eukaryotic methionine salvage pathway.
  Journal
FEBS J 275:4111-20 (2008)
DOI:10.1111/j.1742-4658.2008.06552.x
Reference
PMID:18391471
  Authors
Ashida H, Saito Y, Kojima C, Yokota A
  Title
Enzymatic characterization of 5-methylthioribulose-1-phosphate dehydratase of the methionine salvage pathway in Bacillus subtilis.
  Journal
Biosci Biotechnol Biochem 72:959-67 (2008)
DOI:10.1271/bbb.70651
Reference
PMID:10613873
  Authors
Kitabatake M, So MW, Tumbula DL, Soll D
  Title
Cysteine biosynthesis pathway in the archaeon Methanosarcina barkeri encoded by acquired bacterial genes?
  Journal
J Bacteriol 182:143-5 (2000)
DOI:10.1128/JB.182.1.143-145.2000
Reference
PMID:12644499
  Authors
Mino K, Ishikawa K
  Title
Characterization of a novel thermostable O-acetylserine sulfhydrylase from Aeropyrum pernix K1.
  Journal
J Bacteriol 185:2277-84 (2003)
DOI:10.1128/JB.185.7.2277-2284.2003
Reference
PMID:18520135
  Authors
Tanabe S
  Title
Development of assay methods for endogenous inorganic sulfur compounds and sulfurtransferases and evaluation of the physiological functions of bound sulfur.
  Journal
Yakugaku Zasshi 128:881-900 (2008)
DOI:10.1248/yakushi.128.881
Reference
  Authors
Nishizuka Y, Seyama Y, Ikai A, Ishimura Y, Kawaguchi A (eds).
  Title
[Cellular Functions and Metabolic Maps] (In Japanese)
  Journal
Tokyo Kagaku Dojin (1997)
Reference
PMID:19270684
  Authors
Gutierrez JA, Crowder T, Rinaldo-Matthis A, Ho MC, Almo SC, Schramm VL
  Title
Transition state analogs of 5'-methylthioadenosine nucleosidase disrupt quorum sensing.
  Journal
Nat Chem Biol 5:251-7 (2009)
DOI:10.1038/nchembio.153
Related
pathway
nau00010  Glycolysis / Gluconeogenesis
nau00250  Alanine, aspartate and glutamate metabolism
nau00260  Glycine, serine and threonine metabolism
nau00290  Valine, leucine and isoleucine biosynthesis
nau00430  Taurine and hypotaurine metabolism
nau00480  Glutathione metabolism
nau00620  Pyruvate metabolism
nau00640  Propanoate metabolism
nau00770  Pantothenate and CoA biosynthesis
nau00900  Terpenoid backbone biosynthesis
nau00920  Sulfur metabolism
KO pathway
ko00270   
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